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Phylum Mollusca (molluscus - “soft bodied”) Bilateral, triploblastic, Eucoelomates VERY successful, believed to have 150,000 species… 4.5 million years of evolutionary success… Found in most environments, exploiting most habitats (terrestrial & aquatic) 3 body regions: head/foot, visceral mass, mantle Success due to: Mantle Highly muscular tissue that surrounds the visceral mass (guts!) Many mollusks can extract calcium from water/food, secrete a shell from this tissue Mantle cavity – space between mantle and “foot” (gas exchange, excretion, release of gametes, locomotion) Success due to: Shell Ability to extract calcium from water/food, cement together Secreted by mantle Protection from predators & weather 1 piece – conch, helmit shells, whelks, nautilus 2 piece – clams, oysters, scallops Reduced internal or no shell – slug, squid, cuttlefish, octopus Success due to: Muscular Foot Modified mass of muscle, based on evolutionary use… Digging (clams) Swimming (scallops) Crawling (snails & slugs…mucous gland for land varieties!) Grabbing (squid, octopus, cuttlefish) Communication (squid, octopus, cuttlefish) Success due to: Radula Most have a “rasping tongue” that is used to scrape food off of substrates… Reproduction, Respiraton, Nervous Mostly dioecious (males and females), a few are monoecious (cross fertilization is the rule)… Gills - located between the mantle and the visceral mass (in the mantle cavity) From simple ganglia to most advanced invertebrate brains (great muscle control, incredible sensory control…) Digestion, Excretion, Circulation Great variation, due to habitat Complete digestive tract, with anus Open circulatory system, except cephalopods Class Polyplacophora most primitive; entirely marine; oval bodies with 8 dorsal plates. Broad, flat foot for locomotion Chitons Class Gastropoda “Stomach footed” mollusks most numerous! mostly marine, some freshwater & terrestrial; Visceral bodies enclosed in coiled shell during early development (most retain the shell) Broad, flat foot for locomotion (terrestrial species secrete slime!) Slugs & Snails Class Bivalvia Shell consisting of 2 valves or halves Wedge shaped, muscular foot for digging and locomotion Filter feeders! Sedentary lifestyle – loss of head/radula… Siphons draw in water – passes over gills for respiration & for feeding Clams, oysters, mussels, scallops Class Cephalopoda Marine predators (fast swimmers) “Head footed” mollusks (most advanced!) Reduced, internal shell (or absent!) CLOSED circulatory system Highly developed visual system Tentacles/arms w/suction cups Jet propulsion for movement Squid, octopus, cuttlefish, nautilus Several other smaller, less studied classes