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Transcript
http://www.youtube.com/watch?v=Gg57Id
z7Bs4
 http://www.youtube.com/watch?v=vP_td9
q_344
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General
Name means soft body
this is one of the largest animal phyla,
besides Arthropoda
 There are nearly 75,000 species and
some 35,000 fossil species
 Very diverse group (snails, clams,
octopus)
 Range from simple organisms to some
of the most complex invertebrates
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
Hawaiian Bobtail Squid
Blue Ringed Octopus
Ecological Relationship
Habitats: from tropics to polar region
 ponds, lakes, streams, surf, ocean depths,
etc
 Most live in the sea
 Include herbivores, carnivores, and filter
feeders

Economic Importance
many are food for humans
 produce pearls (most from Eastern
Asia)
 Some molluscs are destructive (Great
Lakes, Snails and Slugs in gardens)

Form and Function
Body Plan –
 consist of head-foot portion (cephalopod)
and a visceral mass (nervous system)
◦ head-foot portion – contains feeding, cephalic
sensory, and locomotion organs
◦ visceral mass – contains digestive, circulatory,
respiratory, and reproductive organs
Form and Function
Body Plan –
 two folds of skin make up a protective
“mantle”
◦ mantle houses the lungs and secretes a shell
in many species
Form and Function
Head-foot –
 many have a well developed head
containing sensory organs
◦ range from simple to complex eyes

Foot is used for locomotion
Form and Function
Head-foot –
 Unique structure inside mouth called “radula”
◦ It is a tonguelike found in all molluscs
except bivalves (shell fish)
◦ Has rows of tiny teeth that point backward
◦ It has about 250,000 teeth
◦ When protruded, it can scrape, pierce, tear,
or cut food material
◦ http://www.youtube.com/watch?v=mLVDwl
rSq5U&list=TLDlxJTSuqf42kySqHbo0jexNd
vt3BkIqS
Form and Function
Head-foot –
 Foot can be adapted for locomotion,
attachment, or a combination of
functions
 Some modifications include – “hatchet
foot” of bivalves, funnel for jet
propulsion in squids and octopi
Form and Function
Visceral Mass –
 (mantle and mantle cavity)
 mantle – the outside of the skin extending
from the visceral hump that runs down each
side of the body, protecting soft parts and
creating a space
Form and Function
Visceral Mass –
 (mantle and mantle cavity)
 mantle cavity – houses respiratory organs
 products from digestive, excretory, and
reproduction systems empty into the mantle
cavity
 in aquatic species, surface cilia create water
currents that continually flush out waste
Form and Function
Visceral Mass –
 Shell (when present) typically has three layers
◦ outer layer
 Composed of a protein called “conchiolin”
 Growth only occurs at the edge of the shell
◦ middle layer
 Composed of densely packed prisms of calcium carbonate in a
protein matrix
◦ Inner layer
 Composed of calcium carbonate sheets laid down over a thin
protein matrix
 This layer is continuously secreted by mantle surface, so it
becomes thicker during the animals life

Calcium for the shell comes from environmental
water, soil, or food
a. Outer layer b1. middle layer
b2. Inner layer
Form and Function
Internal Structure and Function –
 there is an open circulatory system
◦
◦
◦
◦
pumping heart
blood vessels
blood sinuses
closed circulatory system
Form and Function
Internal Structure and Function –
 Digestive tract is highly specialized
according to feeding habits
◦ Most have a pair of kidneys
 Ducts of kidneys in many forms serve to discharge
eggs and sperm
Form and Function
Internal Structure and Function –
 Nervous System consists of several
pairs of ganglia (nerve cells) with
connected nerve cords
 Most molluscs are dioecious
◦ Some are hermaphroditic
Mating Leopard Slugs
https://www.youtube.com/watch?v=FhVi4Z6CjZk
Class Polyplacophora: Chitons
are somewhat flattened and have a surface that
bears eight plates, or valves
 name means “bearing many plates”
 most are between 2 and 5 cm
 commonly occur on rocky surfaces in intertidal
regions

◦ if detached, they can roll up like an Armadillo for
protection

Sexes are separate
Class Gastropoda
(gastro – stomach, podos – foot)
 By far the largest and most diverse class

◦ Contains about 40,000 living species
Includes snails, limpets, slugs, whelks,
conchs, periwinkles, sea slugs, sea hares, sea
butterflies, etc.
 Often sluggish, sedentary animals

◦ Because of heavy shells and slow locomotion
organs
Snails
Snails
Limpets (Owl Limpet)
Limpets
Slugs
Whelks
Whelks (Lightening)
Conchs
Conchs
Conchs
Periwinkles
Periwinkles
Sea slugs
Sea slugs
Sea slugs
Frosted Sea Slug
Sea hares
Sea hare
Sea hare
Sea butterflies
Class Gastropoda

Basically bilaterally symmetrical, but
because of “torsion,” the visceral mass is
asymmetrical
◦ Torsion – a twisting process that occurs
during development
Class Gastropoda

Three subclasses –
◦ Prosobranchia (largest)
 Almost all marine
 Periwinkles, limpets, whelks, conchs, abalones, slipper
shells, oyster borers, rock shells, cowries
◦ Opisthobranchia
 Sea slugs, sea hares, nudibranchs, canoe shells
◦ Pulmonata
 Mostly land and freshwater snails and slugs
 Usually have two pairs of tentacles
Nudibranchs
Nudibranchs
Class Bivalvia
(two-shelled)
 Includes mussels, clams, scallops, oysters,
shipworms
 Most are sedentary
 Unlike gastropods they have no head,
radula, and little cephalization

Class Bivalvia
Most are marine, but many live in
brackish water and in stream, ponds, and
lakes
 Their two shells (valves) are held
together by a “hinge ligament”
 “Adductor muscles” work in opposition
to the hinge ligament and draw the
valves together
 Zebra mussels

Zebra mussels (Lake Mead)
Preventing Mussels in Idaho
https://www.youtube.com/watch?v=J4EVAy8ad
Mk
 http://www.youtube.com/watch?v=T18E58vOTu
s

Class Cephalopoda





most complex species
include squids, octopuses, nautiluses,
cuttlefishes
All are marine
All are active predators
Have an odd body plan
◦ Develops as the embryonic head and foot become
indistinguishable
◦ The ring around the mouth bears arms and
tentacles
Class Cephalopoda

Range from 2 to 3 cm to 13 meters
Giant Squid
Slash marks from Giant Squid
The Search for the Giant Squid
https://www.youtube.com/watch?v=EbubjXsA4S4
Class Cephalopoda

Most have special pigment cells called
“chromatophores” in their skin
◦ By expanding and contracting, they produce color
change
◦ Some color change are for protection to match the
background
◦ Most change is behavioral, associated with alarm or
courtship
◦ Many deep-sea squid are bioluminescent

When the animal is alarmed, it releases a
cloud of ink through the anus to form a
“smokescreen” to confuse an enemy
Squid
Mimic Octopus and others
https://www.youtube.com/watch?v=tLTWFnGmeg&feature=related
Octopuses
Octopuses
Octopuses
Blue Ringed Octopus

The most venous octopus.
◦ This small mollusc lives in warm, shallow reefs off the coast of Australia,
new Guinea, Indonesia and the Philippines.
◦ It has a life span of about one and a half years.
Blue Ringed Octopus
◦ It is said that the venom of this octopus could kill 26 adults
in just a few minutes.
◦ There is no antivenin for treatment.
◦ Fortunately, these octopuses do not attack humans.
◦ Injury typically occurs when a blue-ringed octopus is
stepped on or picked up.
Nautiluses
Nautiluses
Nautiluses
Cuttlefishes
Cuttlefishes
Cuttlefishes