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The Cell: Basic Unit of Life Stephen J. Salek Prokaryotes and the Evolution of Eukaryotes Early earth and the origin of life • Universe is 10-20 billion years old • Atmosphere H first then other gases • Earth solidifies 4.1 bya • 1st photosynthetic prokaryotes 3.5 bya Universal Cell Components • • • • cell membrane cytoplasm genetic material (DNA) ribosomes – for protein synthesis Campbell Fig. 7.4 Bacteria The Smallest, Simplest Cells Campbell Fig. 27.2 rod-shaped spherical ALL BACTERIA ARE PROKARYOTES. spirilliform Prokaryotic Features • DNA is in a single-stranded loop – No chromosomes, mitosis, or meiosis – Lack a membrane-bound nucleus • No mitochondria, chloroplasts, or other organelles • Cell walls are chemically unlike any eukaryote cell walls Bacterial Cell Features Campbell Fig. 7.4 Advanced Prokaryote Features • interior membranes • bacterial flagellum • some are photo-autotrophic compare Campbell Fig. 27.9 Campbell Fig. 27.6 Campbell Fig. 27.5 Bacteria are Important • Pathogens (“disease-causing”) are parasites on living cells and organisms • Decomposers rot dead material, recycling essential nutrients • Mutualists with – plants: root-knot rhizobia fix nitrogen – fungi: cyanobacteria are part of lichens – animals: intestinal bacteria digest cellulose Commercial Bacteria • Lactose-users make yogurt and cheese • Soil bacteria make many antibiotics • Intestinal bacteria used as “lab rats” in cell biology • Cyanobacteria are important for marine and freshwater food chains 3 Domains of Life Campbell Fig. 27.1, 27.11, Table 27.2 • Archaea are also prokaryotes • Recently separated from Bacteria – Cell wall chemistries are different – Archaea genes more like eukaryotes’ • Only Eukarya have a proper nucleus Phylogeny of Domains prokaryotes Bacteria Archaea Eukarya Archaea • unusual environmental tolerances: – extreme halophiles – extreme thermophiles – methanogens Campbell Fig. 27.10 Eukaryotes • true cell nucleus: – nuclear membrane – linear DNA, organized into chromosomes – replicate by mitosis and meiosis • Plants, animals, and fungi are eukaryotes. • Many unicellular algae and protozoans are also eukaryotes. Evolution of Eukaryotic Cells Campbell Fig. 28.2 Step 1: Nuclear and endoplasmic membranes (kingdom Archezoa) Step 2: Organelles acquired by symbiosis (remaining 7 kingdoms) Organelles • Bacteria that were parasites, became mutualists – mitochondria burn sugars, etc., for energy – chloroplasts photosynthesize more energy – chloroplasts were lost during evolution of many living eukaryotes, such as animals and fungi Mitochondrion Campbell Fig. 7.17 External and internal membranes (compare to early stages of eukaryote cell evolution) Chloroplast Campbell Fig. 7.18