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Modern Ideas on the Origin of Life copyright cmassengale 1 Formation of the Earth • 4.5 billion years ago • Cosmic debris attracted to one another • Collisions = heat = melting • Atmosphere = hydrogen cyanide, CO2, CO, N2, H2S, H2O Formation of the Earth • 3.8 billion years ago • Cooling = oceans • Brown oceans due to dissolved iron • Life Begins – Prokaryotic, anaerobic, heterotrophs First Organic Molecules • 1920s – Oparin and Haldane • Earth’s early atmosphere had all elements necessary to cause reactions producing organic molecules (amino acids) • Gases, UV light, lightning PRIMORDIAL SOUP HYPOTHESIS First Organic Molecules • 1950s – Miller and Urey • Suggests how mixtures of organic compounds necessary for life could have come from simpler compounds present on primitive Earth First Organic Molecules Animation of Miller and Urey’s Experiment • http://ircamera.as.arizona.edu/NatSci102/ movies/anim_miller_urey_exp.gif • Proved amino acids could be produced in primitive Earth’s conditions Early Life - the first cells were probably heterotrophs that simply absorbed nutrients and ATP from the environment. - as these substances became rare, there was strong selection for cells that could manufacture their own energy storage molecules. Early Life - the second type of cells were probably like green-sulphur bacteria which in the presence of sunlight, are able to photosynthesize. Early Life - the evolution of oxygenic photosynthesis was MAJOR. It allowed life to exploit more habitats. These stromatolites, which date to > 3 bya are microbial communities. Early Life - about 2.3-1.8 bya, the concentration of oxygen began to increase in the ocean. Oxidized iron shows rust bands in sediment. Endosymbiont Theory • 1966- Lynn Margulis • Explains origins of mitochondria and chloroplasts • Mitochondria and chloroplasts were once free-living bacteria which had a mutualistic relationship with another bacterium Endosymbiont Theory • Mitochondria and chloroplasts are same size as bacteria and contain similar structures (DNA, ribosomes) and reproduce by binary fission infolding of membrane Endosymbiosis - mitochondria and chloroplasts (Margulis - 1970's) Evolutionary Tree – shows evolutionary pathways of living organisms So, reconstructing the patterns of relatedness among these ancient life forms is difficult. Different genes give different patterns of relatedness among domains