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New
species
of
Streblus
E.J.H.
and Ficus
(Moraceae)
Corner
of
Botany School, University
Cambridge,
U.K.
Summary
New
Taxa.
novahibernica
Notes
F.
are
Isl.);
F.
and
given
on
Streblus
nov.
F.
and
F.
the
Solomon
S.
subcuneata
solomonensis,
with
key
a
New
Ireland,
Ficus
its
to
ascendens
Isl.);
F.
hesperia
(sect. Adenosperma,
nov.
sp.
S.
single species
sp.
nov.
Caledonia).
New
lapidaria
New
Sycocarpus,
pendulinus,
and
with
nov.,
(subgen. Pharmacosycea,
nov.
sp.
(sect.
cryptosyce
adenosperma (Rotuma),
var.
servula
F.
sect.
(sect. Paratrophis,
nou.
sp.
malaitana
var.
Solomon
Protostreblus,
sect.
sclerophyllus
S.
Ficus cristobalensis
Sycidium,
Lour.
Streblus
—
(Solomon Isl.);
(sect.
nov.
sp.
New
Guinea);
F.
Guinea).
illiberalis,
subtrinervia
F.
(Solomon Isl.),
allies.
Streblus Lour.
sect.
Folia
spiraliter
laminam
Protostreblus
lamina
disposita;
ovata
nulla.
duplicatis. Cystolitha
S.
Typus:
—
The structural peculiarity of this
new
of Streblus. The lax
Artocarpus,
(Corner,
as
the
Maclura, and
1962), the
of Streblus.
The
arrangement,
e.g.
basal veins
may
occur
spiral
F.
sycomorus,
veining
in
S.
Paratrocon-
and
growth
section appears
the
as
F.
occurs
variegata,
pendulitius, with
be
a
albipila,
S.
to
the
Ficus,
understand them
remnant
a
in
contrast
of the
ancestry
longer lamina,
fits
of the shapes of leptocauly with
elongatus
which
I
as
the distichous is manifest
to
vestige of
one
F.
in Streblus
Moraceae, such
develops applanate,
to
subcordate
clearly antecedent
in which
sense
ovate
intercostals is unknown in
In various
arrangement
sections of Ficus where it is
most
Similar intercostal
profusa.
the broad
in
new
mediam
subincrassatis
The
Paratrophis.
transverse
genus.
with acuminate tip,
leaf-shape
in
of the
horizontal in its
foliage. Thus this
parallel examples
spiral
more
sect.
numerous
rest
Broussonetia
in
Insulis Solomonensibus.
ascendens,
arrangement of the leaves is
transition from the
twig becomes
with ascending,
spiral
also in the
prevails
ut
section lies in the combination of the Moras-like
lamina with prominent basal veins and
rest
ad
sect.
basalibus
foliaceis
cotyledonibus
reproductive characters of Streblus
distichous which
costis
subcordata,
v.
phis; embryo radicula incumbenti elongata,
the
nov.
intercostis transversalibus numerosis. Inflorescentia
elongatis,
leaf with the
sect.
grossularioides, and
F.
F.
somewhat prominent
and
ascendens
may
have its
nearest
relation.
The
second
I have
1962,
p.
several
peculiarity
pointed
is
p.
40—53).
occurrence
it would be
which
bridges
primitive
of which is
to
have
expected
near
at
the
of the
one
nature
of Streblus
mainly
come
If S. ascendens is
domain of S. pendulinus and
embryo of
the
relatively
of Ficus seemed
1967,
exactly where
the
the distribution
221),
groups
(Corner,
out
a
centre
in the Solomon Islands.
sect.
Paratrophis (Corner,
Melanesian, and have shown how
from
relic
species
the Jurassic
Melanesian Foreland
of the ancestry
of distribution of
that of the New Caledonian S.
of Streblus, it
sect.
Paratrophis
occurs
in the
sclerophyllus (p. 399), the
the difference between those of S. ascendens and S.
pendulinus.
BLUMEA
394
Figure
X
7;
d.
I.
l.
Streblus ascendens.
s.
and
t.s.
ripe
a.
fruit
Twig,
X
VOL.
½;
(endocarp
b.
t.s.
XVIII,
female
and hilar
No.
spike,
2,
X
1970
5;
plug hatched),
c.
X
female flowers and
10.
(BSIP 8624).
young
fruits
E.
There
of
are,
course,
S. ascendens has been
which
I add
note
a
found, and
that it is
on
sections of
as
the
recent
as
stenieae)
is much closer
Sloetia,
and Pseudotrophis
tion,
to
and
several species
Streblus
e.g.
genera
phyllus with
of S.
ways.
1 —8
with
pass
Ficus
(Moreae) than
more
over
flowers
1—9
state
ancient
a
to
But
not
of Streblus, in
has
from
pachycaul
same
in the wide
to
represented by
yet
with
comprehensive
ascendens,
Arbor —23
a
are
merely
are
sense
given
as
a
are
new
so
in
common
very
two
genera.
The
fifth
easily spread
(Ficeae of Hutchinson,
satisfactorily
defined in Hut-
in bud and the
stipules
prime marks of the subfamily.
as
The
as
be found in
their vegetative characters
of advanced
collection
a
(Taxo-
many,
must
the
—9
has been missed. These
spirally folded
not
disappeared.
sp.
m
as
of
species
an
section Protostreblus endorses
much of the previous evolutionProtostreblus is the relic
sect.
puberuli,
mm
mm
saepe
sprinkling
assist
of the
to
though these
the present
state
may
of
have
Streblus,
of species with small distichous leaves of similar
the understanding
con-
—
trunco
Figure
longo, basi rotundata
1.2
2—3
subcoriacea,
v.
and Sloetia prevent.
I.
ambitu
sparsim. Ramuli
and phyletic classification of plants which
Pseudomorus,
longae caducae puberulae.
subdenticulata,
Dorstenia,
or
Streblus. Ficus, reduced
Pseudostreblus,
nou.
alta,
a
as
griseo; latice albido copioso; foliis
—15
—6 flowers
2
macro-
Teonongia without mention
as
in
S.
considerable diversity in detail of flowers, seeds, and embryo. Thus
genera
such
microgenera,
—6
flowers, and
2
distinc-
with
pendulinus
One could much more
Sparratosyce
distichous leptocaul,
beginnings
vergence,
v.
2 —6
sense,
Streblus,
in
immediateconnection with Fatoua, Morus,
from the
would be
S.
S.
arrangement of the leaves inherited from the extinct stock of Streblus. The section
no
come
my
Artocarpoideae
which
always amplexicaul,
the inflorescence.
on
the chief distinction between these
even
stock of Moraceae that has
states
spiral
with
which is
Diplothorax
to use
the exclusion of
scheme).
(Dor-
whereas
Streblus,
as
reduction from that with
this view. By analogy with Ficus, which has retained
ary
What I
Sterile, Pseudostreblus,
Dorstenia.
zeylanicus with
S.
with few flowers is
monoica which breaks
regard modem Streblus
I
ilicifolius
S.
Moreae), and
(Pseudotrophis,
four tribes
means
accords
tree
The affinity of Pseudo-
(Paratrophis, Moreae),
Moreae),
fruit, seed, and embryo
my
of
Moreae, Strebleae, Dorste-
than four female flowers
sharp distinction from Fatoua and Dorstenia.
Olmedieae of
no
large
number of female flowers defies such
chinson's classification; the leaves of many
by
a
plant
of its sapling
three distinctions. In Hutchinson's scheme Moreae
place Bleekrodea (Fatoueae) with Sloetia
in
male
status.
Taxotrophis
and
Paratrophis
are
to
misinterpreted by Gagnepain, and he recognizes
var.
asper
indicate,
relict
no
nature
rather than with Fatoua, and Sloetia
points
in which the
Hutchinson continues
asper,
characteristic
points
As yet
learn the
other genera, and tribal, generic, and sectional characters but
so many
Streblus
to
and confusing rendering.
Pseudotrophis
(Pseudomorus,
trophis, Strebleae). The
other
interesting
395
species besides that of leaf-shape;
cotyledons.
four tribes (Fatoueae,
among
artificial
glaber with
flowers
16
even
(Moraceae)
classification of Moraceae by Hutchinson (1967).
differ from Strebleae mainly in having
are
two
accords with its
rare
sterile Fatoua, Dorstenia, and Morus
There
flat
and Ficus
lobate leaves. That it becomes
even,
is with Sloetia in all
(Fatoueae)
streblus
and thick
it will be
clearly
the previous,
is
Streblus
species of
Streblus, defined on the inflorescence, female flower, fruit, and embryo,
Hutchinson distributes
nieae). This
New
fruiting tepals
have dentate and,
may
with its lamina;
treat
Corner:
other differences between these
has short
pendulinus
S.
H.
J.
m
attingenti,
laxe
mm
9
—
alata;
cortice fuscibrunneo
dispositis. Ramuli foliaque albidi-
glabrescentes ochraceibrunnei. Stipulae
crassi,
Lamina
subcordata,
superne
ad basim
spiraliter
:4
X
5—8
etiam fere
scabriuscula,
sicca
cm,
ovato-acuminata
subpeltata, integra
superne
v.
apice
distanter
grisea subtus brunnea;
BLUMEA
396
Figure
2.
Streblus
(Queensland);
c.
St.
pendulinus.
John
and
Leaves
Fosberg
VOL.
of various
15271
(Rapa);
XVIII,
No.
collections,
x
d. Stone 5194
2,
½;
1970
a.
f.
x
926
10;
Vickery 3507 (N.S.W.), male with bracts, X 7; g. Kajeswki
e.
X
4,
flowers
and
young
fruit
X
6.
Forbes
(Rota);
e.
797K (Hawaii);
NGF
b.
Mueller
26170 (Papua). Flowers;
(New Hebrides), inflorescence,
E.
H.
J.
Corner:
New
species of
costis lateralibus utrinsecus 3 —5, subtus
transversis;
basalibus
nullis;
solitariae
basim
costis
2 —6
petiolo
2
2
longus,
1 —1.3
mm
fructu4 —4.5
ovario
mm
—22
mm
pedunculo
pendulinus
I
give
some
4—4.5
(Endl.)
X
obtusis
3
v.
mm,
M.;
1.5 —2
reduced Strebleae
solomonensis
Figure
and
was
to
Corner
collected
on
of
embryo,
x
bracteis
Flares
liberis
longis; stylo
primum
feminei sessiles;
subaequalibus,
subnullo. Fructus
drupaceus,
nulla.
Kolombangara
Corner
(1962)
the 2-flowered
Moreae and Pseudomorus
(1962)
224.
—
several occasions
3. Streblus solomonensis.
end-views
puberulo;
axillares
puberulae,
in
pr.
222.
Inflorescentia
a.
4.
Fruit in l.s.
(RSS 65).
mascula ?
—
Figure
Paratrophis
seen
Figure
3.
during
the
as
mature
a
species. According
26170 should belong
a
embryo of
b. pyrene
in
hilar
in
Therefore,
section of
Royal Society
(endocarp hatched);
silvis
2.
926 in Pseudomorus (Moreae).
to
convo-
In
Kokove. Typus: BSIP 8624 (L).
of NGF
spike
mm
7
maturitate aurantiacus
brunneo duro tenui. F.mbryo cotyledonibus
tribal distinction is less than specific. I have not
This
mm,
puberulis
mm
Cystolitha
Taxotrophis (Strebleae) and that of Kajewski
S.
1.5
glandulis
femineae spicatae
illustrations of this widespread, yet insufficiently known,
Hutchinson's classification,
to
subtus leniter elevatis,
attingentibus,
unilateraliter praeditae,
tepala projiciens,
incumbenti.
F.
397
interioribus paulo majoribus, fructum longinque includentibus;
tandem ultra
radicula elongata
X
7 —12
longo, stigmatibus duobus
obovoidcus,
4 —9,
latis, dein lateraliter extensis, puberulis.
locis planis, Insulis Solomonensibus,
S.
gracili. Inflorescentiae
mm,
(Moraceae)
ad mediam laminam
obovato-rotundis,
et
intercostis
elevatis;
1,
and Ficus
multiflorae, sulco sterili
mm
longis,
longis
(? ruber); endocarpio
lutis,
15
maturescentes;
versus
4,
utrinsecus
longae,
cm
semicircularibus
tepalis
basalibus
Streblus
Streblus;
S.
pendulinus.
Expedition
view;
c.
I
the
l.s.
to
the
embryo
BLUMEA
398
Figure
fruit
(l.s.
4. Streblus
with
sclerophyllus.
a.
Twig
endocarp hatched),J.
M.
VOL.
with
male
Veillon
XVIII, No.
spike,J.
1296
,
X
2,
1970
M. Veillon
5.
116,
x
½;
b.
female
spike
and
c.
E.
Solomon Islands.
Corner:
H.
J.
Additional
latex white, copious; lamina
4—8
long and fertile
mm
wide, the tepals
mm
1.5
subglobose,
(living),
sclerophyllus,
Frutex
Ramuli
points
—20 cm
v.
arbor
part
4 —7 cm
with red
7
—
alta, inermis,
elliptica
caducae
longae,
mm
subobovata, apice
v.
—15
bark
with
plug; frequent
reticularis supeme
I,
Figure
—
—13
1
alabastro
mm
mm
2
longis,
7
m
1900
alt.
2
X
11 -5
—
foliis distichis.
crassi,
mm
fuscibrunnei.
brunneola;
v.
intercostis
impressis;
anguste
3 —4 cm,
subrigida
costis lateralibus
laxis, vcnulis
I—2
costis basalibus utrinsecus
gracili. Inflorescentiae spicatae simplices axillares solitariae;
denticulatis; femineae pedunculo
internis 1.5—1.8
mm;
Fructus
io
stigmatibus
crassiusculis
cotyledonibus
montanis Novae
Caledoniae,
4,
8 mm,
X
stigmatibus persistentibus, tepalis
accumbenti,
silvis
supeme
folded
—20
X
1.5
mm,
sulco sterili praeditae. Flores feminci tepalis 4 subglabris,
—7 cm,
pistillodio quadrato puberulo.
crassatus,
to
up
basi anguste cuneata,
longo acuminata,
lati, tepalis puberulis; staminibus
mm
forest
wide
mm
variously
longa puberula, floribus paucis (c. 5) praeditae; masculae pedunculo
parte fertili
mm,
externis
—12
10—11
and
4.
Ramuli
glabrescentes.
mm
the
in
lanceolatae. Lamina
ovato-reniformibus peltatis
fertili
parte
15—22 mm,
leaf-scars;
persistent
white; fruits
impressis, subtus planis, areolis brumieolis;
brevibus; petiolo
bracteis
399
videtur dioica; latice flavidula;
ut
subtus len'ter elcvatis,
7 —10,
with
conduplicate
coriacea, marginc incurvo subdentato, levis, sicca griseiviridis
utrinsecus
(Moraceae)
grey,
the anthers
dots,
embryo
petiohque primum puberuli,
Stipulae
and Ficus
long; bracts reniform-peltate; male flower-buds
(sect. Paratrophis).
nov.
sp.
—6 m
these:
are
green
purple-black;
Streblus
long; male spikes solitary, axillary, with penduncle
the hilum without sclerotic
cotyledons,
S.
species of
New
mm
drupaceus ruber,
subaequalibus,
plicatis.
m
in
longis, incurvatis;
basim
subin-
versus
Semina radicula
paulo amplificatis.
Boulinde 950—1100
mt.
longis. Flores masculi
2 mm
filamentis —3
Cystolitha
elongata
nulla.
alt. Typus: J. M.
In
Veillon
1296 (P).
Collections:
This is
upperside.
seems
J.
Veillon
July
26
1296,
The reniform bracts
abied but the
scant
but it
denticulate leaf of S.
(type, P); J.
coriaceous
place
it in the
cotyledons
are
unlikely
seems
female
1967,
by the stiffly
distinguished
closely
material is
M.
M. Veillon
leaf with
conduplicate
26
April 1964,
the veins impressed
of S. pendulinus
complex
that the species
116,
much
as
male.
on
the
with which it
S. solomonensis. The
in
develops the larger and closely
pendulinus.
FICUS Linn.
The
species here described
cbeck-bst
mij
have been
(Corner, 1965).
This
given
places
the number with which I hisert them into
them in natural abiance and
aration of the bsts
of identifications, sometimes running
which I have had
to
Not much that is
in the
or
prep-
thousands,
prepare.
new
comes now
from
western
Malesia. New Guinea, New Britain,
and New Ireland have continual interest and, indeed,
300
helps
into hundreds
the Solomon Islands. Among
some
cobections made by the Forest Department, BSIP, since the Royal Society Expedition
in 1965, I found the new
very
species
of
sect.
Sycidium, namely
well developed in the Solomons, and the interesting
cosycea,
namely
However, it is
appear.
F. cristobalensis
in
the
sections
var.
malaitana;
Adenosperma
it
may
F.
new
web
hesperia, this
variety
prove
and Sycocarpus
to
in
be
section
being
subgen. Pharma-
a
distinct species.
that novelties
When I revised recently Adenosperma and added die new species F.
continue
to
pilulifera and
BLUMEA
400
F.
suffruticosa,
1969).
I
servula, which help
known F. subcuneata. This
expecting
bridge
to
species
suffruticosa
to
is
light
a
nearest
remarkable
to
tree
bearing
on
between F.
gap
new
fig
but it is
F. arbuscula in
keyed
out on p.
differences
405, yet the
bridged by further
collections. It is
The alliance confirms
my
sympatrically
in
the
to
to
are
be
study
such
hoped
as to
that
parent
species
well
had been
F. subcuneata is
my
a
tall
small shrub. F. lapidaria is
into
the three
F. servula is
new
species
may now be
that they
a
are
may
as
be
studied ecologically.
that evolution has
through
a
duplicates and distributed
one to suppose
general conclusion for Adenosperma,
domain of the
now
I
Though readily distinguished,
lead
they
(Corner,
however, have exceeded
a
up
the
Hypogenae and
Amphigenae; and
ser.
them.
alliance
species, F. lapidaria
suffruticosa and
lapidaria. Unfortunately,
the world before it has been possible
knit
other
two
ser.
species,
known from single gatherings and these have been split
over
closely
ecological complex.
new
it in leaf and
the ancestry of F.
1970
to add
rather isolated in
was
rheophyte comparable with the bushy
small
the
2,
in this
oblige me
the discovery of allies. The three
hopes and brought
F.
No.
XVIII,
thought finality had been reached
More collections from New Guinea
and F.
tree.
VOL.
structural
proceeded
and ecological
diversity.
Nevertheless, of
more
thorough
Figure
5.
Ficus
all the countries of Malesia Celebes
investigation
cristobalensis
var.
of its
Ficus
malaitana.
Fig.
seems to
be that
most
flora.
x
2;
flowers,
x
10.
BSIP
10568.
in need of
E.
H.
J.
113B. F. cristobalensis
var.
Differt
a
typo
—6
in omnibus
mm
crassi.
lateralibus utrinsecus
14
mm
401
In
silvis
BSIP
—2
cm
intercostis
—3
mm
Tepala
primariis
6. Ficus
smithii).
Auki
10568
,
Malaita, Nazareth,
Figure
F.
and
Plait
(Moraceae)
(subgen Pharmacosycea)
b.
401
var.
malaitana,
et
Lamina
petiolo
2—4;
longo;
3
longae.
11 —23
—-18
9
mm
X
Arbor
(vel 4), libera
v.
plus
Insulis
minus
m
mm
longis;
alta.
costis
cm;
longo. Receptacula
bracteis basalibus 1.5 —2.5
secundariis,
—7
4.5 —7.5
13
—
cellulis
gamophylla. Cystolitha
Solomonensibus,
Malaita. Typus:
10568 (L).
Collections:
(?
2
in pariete nullis.
hypogena.
East
(1967) 78
Strebhis
partibus minoribus, tepalis paucioribus.
Stipulae
10 —13;
lata; pcdunculo
scleroticis
BSIP
Corner
species of
Figure 5.
nov.
Ramuli 5
New
Corner:
30
illiberalis.
m
area,
alt.,
Leaves,
Esisiki,
bark
X
½;
dark
a.
300
m
alt.,
brown,
BSIP 8500
bark
latex
dark
sticky.
(typical);
brown,
latex
copious;
BSIP
1066g,
‘Malifu’ (Kwara'ae).
b. BSIP
7060 (Vanikoro);
c.
BSIP
7844
402
BLUMEA
VOL.
XVIII,
No.
2,
1970
E.
This
agrees
At first I
both
closely with
leaves
larger
even
probably be infertile.
which
entity
F.
to
The
edelfeltii.
F.
may
stands
wassa
be
to
prove
F. copiosa
collections
Such small
trees.
in
the
collections
two
species with
in the
distributed in Guadalcanal ( BSIP 2785 and
the
size which
F.
112A.
it
needed
BSIP
7060,
BSIP
Kolombangara,
to
F. smithii
the
10163,
malaitana
on var.
(1967)
Isl.,
Vanikoro
Sta.
7844,
Vila
south east,
in the
which,
(subgen.
80
Santa Cruz
Ysabel,
R.,
These collections have shown
close
as
to
the persistent
fig-wall, and
is
large
a
tree.
former mixed with F,
discover its sapling and
to
to
novae-
ascertain
reach.
illiberalis Corner
Collections:
8500,
are
may
‘malifu’ (Kwara'ae);
stand
to
been found in Malaita but recent collections show that it is widely
not
georgiae). Field studies
would
much
very
seems
edelfeltii which lacks
stipules and cordate leaf-base, has copious sclerotic cells
F. cristobalensis has
F. smithii
as
but
distinct
a
represent
leptocaul habit,
more
F.
as
to
ways.
trees,
cristobalensis would
of F.
trees
subgen. Pharmacosycea,
distributed
were
403
fruiting twigs and they
their massive
on
I consider
separate
a
and,
(Moraceae)
side-branches of large
represented merely
than normal
Therefore,
and Ficus
cristobalensis but differs in being smaller in all
F.
from small
come
Streblus
species of
New
that the collections
thought
similar and
are
have
very
Corner:
H.
J.
March
group,
north west,
Qarangao,
Dec.
10 m
1967,
that F.
me
Pharmacosycea).
is
illiberalis, which
is
Solomons,
small slender
a
‘bubulia’
1966,
‘bubulia’
tree,
6.
‘nganingaio’ (Nambalua name),
1965,
Oct.
Figure
—
(Kwara'ae);
BSIP
(Kwara'ae).
a
large buttressed
BSIP 8300 is
tree.
tree, is
typical
F.
illiberalis with coriaceous spathulate leaf. BSIP 70601s the second collection from Vanikoro
with
larger leaf with
more
smithii;
as
a
119b.
BSIP 7844
ridge-top
suggests F. smithii.
Collection:
This is
BSIP
a
6602,
Sta.
seems
trees
of F.
doormaniana
var.
Ysabel,
north west,
variety, though I referred it
(Diels)
Binusa,
8
and its
F.
to
pachystemon
male flowers
disperse
of F. pachystemon
nervia has
tree in
m
Fruiting
because of its
(Comer,
1967,
Arbor
cortice
hesperia,
—35
m
lanceolatae,
tundata,
Figure
alta
fuscibrunneo;
scabridipuberuli.
apicem
in the Solomons has yet
of both
to
nov.
(sect.
(subgen.
albido;
1 —1.5
Lamina
Sycidium).
foliis
mm
5 —13
subcoriacea
hesperia. Twig,
x
½; female
two
be
mm
long, but
it has
also be this
may
larger basal bracts (3.5 —5
I have
are
rather
In
few
a
occur-
subtri-
New Guinea F.
disperse males
scarce
and
it
is
mm
been
never
species, I think that the
proved.
there
are
Figure
—
3.2
m
in
some
difficult
to
7.
attingenti, ad
fuscibrunnei.
2—4-5
attenuata,
duriuscula
2
distichis. Ramuli folia
crassi,
x
Pharmacosycea).
forest, figs red, Jan. 1966.
mangrove
83). Though
p.
acuminate leaf
acuminate lamina.
from F. pachystemon.
ambitu trunci
umbrosa,
latice
species
subtrinervia
elongatum gradatim
integra
Fitus
sp.
Ramuli
caducae.
acutum
7.
specimens
leaf alone F.
distinguish by
329A. F.
shortly
collection BSIP 4333
generally only ostiolar male flowers, but
collections.
a
Corner
satisfied with the absolute distinction between these
rence
but the shortly
illiberalis
illiberalis have
that the previous
I think
distinct,
a
be intermediate between F. illiberalis and F.
with F.
agrees
young
to
characteristic collection. It has the small basal bracts
disperse male flowers.
long)
it
tree
Possibly
subtrinervia
F.
subcordate base; it is fertile and suggests that this is
attenuate
easterly variety.
scabrida,
flowers
and
Stipulae
basim
alata
receptacula
3 —4
ovato-lanceolata
cm
basi
et
>
mm
seeds,
grisciviridis;
x
10.
(R.
costis
Schodde
v.
vix,
longae,
symmetrica
cuneata, saepe laticuneata
sicca
v.
minute
ad
subro-
latcralibus
4073).
BLUMEA
404
utrinsecus
4—6(—7)
glandulis basalibus; pctiolo
sine
XVIII,
subtus leniter
ascendentibus,
costis basalibus utrinsecus
elevatis;
vix
VOL.
haud
1,
mm, ad
5 —15
No.
2,
longis,
bracteis
subacutis;
ovatis
v.
bracteis parvis
lateralibus, ostiolo
albidis; cellulis scleroticis
albidis
glabris
Insulis Solomonensibus. Typus:
R.
Collections:
Jan.-Feb.
This lofty
Schodde
tree
It differs in the
may
copious latex.
It
In the Solomons
of the
pedicels but
and F.
flower
out next
in the
a
seem
to
on
but these
a
lenticuli-
mm,
silvis,
In
(L).
8373, 8899, Kolombangara,
7553,
long-attenuate
apex
and fewer
in the
tepals, and evidently
Sycidium
chrysochaete (Corner,
F.
collectors.
melinocarpa by previous
leaf with
venation of the
the
larger
1967,
Solomons,
tree
it
be
may
setose
in
more
pseudowassa
and
to
comes
near
and both of these species
the
lateral
veins and
may
ampelas
have smooth
a
collar, and glabrous
intercostals,
larger,
a
but is has
there is the problem of the large
these F. hesperia
glabrous
F.
soronensis. F. schumanniana has
var.
longer
base and basal glands,
cuneate
in
102),
89,
p.
to
more
Scabrae.
ser.
leaf, the hairiness,
intercostals, basal bracts rarely
the petiole. Nevertheless,
in F.
occur
BSIP
narrowly elliptic leaf with narrowly
acuminate and denticulate leaf with
dorsal gland
to
shape and
clearly
it is
basal veins, lax
pedicels,
I
amphigcna.
relate these species of sect.
to
tepals and style. Outside
schumanniana, but
elongate
Semina
v.
spathulatis,
v.
the lack of sclerotic cells in the fig-wall, and the flowers with
fig,
seeds. F. ampelas has
more
difficult
ovate
in the white
flower-pedicels,
setose
flora, this keys
setose
feminei sessiles
Flores
Cystolitha
sine
copiosis,
mm,
lanceolatis
5,
1—1.5
Solomon Islands.
(type, L);
well have been mistaken for F.
which differs considerably
peduncle
tepalis
valleys,
Buin
—0.8
internis
3,
vivo) globoso,
mm,
(Kwara'ae).
exceedingly
is
setis
masculi ?.
swampy
symmetric, harshly scabrid,
lateral veins, in the
2—5,
videtur
ut
bracteis basalibus
Schodde 4073. Bougainville
Bougainville,near
4073,
‘samotasubi’
1968,
R.
lowland forest of flat
Habitat: in
mm;
(12 —14
sessili; stylo simplice glabro.
minute punctata. Flores
subcarinata,
formia
occluso;
minus dense setoso;
ovario albido
liberis;
lato
receptaculi feminei copiosis.
in pariete
pedicellati, pedicello albido plus
transversis
—\ ( —J) laminae,
apicem dorsalem subglandulosa. Receptacula
mm
9 —12
corpore
intercostis
elevatis;
elongatis
axillaria solitaria, maturitate aurantiaca; pedunculo 4—6
mm
1970
trees
be related (Comer,
the
which
1967,
106).
p.
386. F.
adenosperma
Among
Bishop
H.
St.
19709
some
Miq.
older collections from Rotuma Island,
Museum in
Honolulu,
They
had gall-insects. There is
most
easterly
no
occurrence
District),
from
were
One collection
slopes and ridges.
and
Itutiu
John 19082 (Kilinga,
(Solkope Isl.).
I found four collections of
trees
(19082)
recently
typical
19603 (Solnahu Isl.),
7
—10
m
high,
to
me
from the
adenosperma,
of F. adenosperma
namely
19639 (Uea Isl.),
growing
on
and
the wooded
had seeds with embryos and another
doubt, therefore, that these collections
relates Rotuma phytogeographically
sent
F.
(19603)
natural
represent the
and of the whole section. The discovery
with the Santa
Cruz
Islands,
the Solomon
Islands,
and the New Hebrides, and it emphasizes the separation of Fiji, Tonga, and New Caledonia where
sect.
represented
obliqua,
F.
Adenosperma
F.
prolixa,
does
F.
not
scabra,
occur.
The other collections from Rotuma
and F. tinctoria
ssp. tinctoria,
as
they
New Hebrides.
389. F.
I
subcuneata
provide
a
key
to
Miq. (sect.
this
Adenosperma).
and its
three allies,
as
described subsequently.
occur
in the
E.
H.
J.
Corner:
KEY
i.
TO
Twigs, petioles,
and underside
with
5—25
peduncles
2.
Tree
2.
Shrub
—30
1
basal
3.
Basal
high.
1. Appressedly hairy,
long;
Basal
high.
m
m
12 mm
wide
Fig
25—30
mm
fine
close
—3.5
OF
F.
strigoso-villous
bracts
6—8
bracts
bracts
persistent,
Fig
COMPLEX
of the leaf
basal
Streblus
species of
and
1
(dried);
—3.5
Ficus
(Moraceae)
405
SUBCUNEATA
with
hairs.
spreading
caducous.
Stipules
F.
subcuneata
F.
suffruticosa
long
mm
long
mm
Stipules persistent.
Peduncles
wide
internal bristles rather sparse. Leaf with
intercostals.
suffruticosa
389A. F.
389.B. F. servula,
Arbor
attenuate
apex.
Small
nou.
paginam laminae inferam)
3
crassi.
mm
Lamina
tree —10 m
pilis
17—22
4-8 —7-8
X
costis basalibus utrinsecus
4—7;
Receptacula
Receptacula
axillaria
bosis;
setis internis
1—2
NGF
Typus:
This
agrees
stipules
smaller basal bracts
from F.
or
more
tepals
as
as
petiolo
puberulis,
ligulato-lanceolatis,
montis
10—20
mm.
0.5—1.5
persistentibus;
gib-
6—8
superantibus,
ovarium
laterali. Semina
(in silva?),
sicca
intercostis
cellulis scleroticis in pariete
subsparsis;
Nova Guinea,
subcuneata but the hairs
the
veins
much
are
straight hairs
particularly
well
2;
hirsutae.
nitida, levis,
superne
appresse
are
ut
liberis;
in F. subcu-
Bulolo,
Morobe
F.
the leaf is
appressed,
less prominent
are
and
largely
are
seems to
trichocerasa
acuminate leaf with
are
A
of F. subcuneata
(lateral bracts
glabristipula
longer
slight undercoat of
the complex
connect
var.
petiole
and
the
the much
Thus it differs also
sparse.
slightly larger.
more
glabrous,
smaller, subsessile, and shortly hairy,
persistent, and the internal bristles
less
(ad
subacuto
apice
hirsuta, glabrescentia; pedunculo
longis,
mm
the basal bracts of which
trichocerasa,
F.
shorter
are
suffruticosa,
microscopic,
with
with F.
persistent, the figs
are
media
appresse
subobovata,
(—13), subtus valde elevatis;
rubribrunneo; stylo
jugum
lapidaria
(CGE).
acuminate,
not
v.
subcoriacea integra,
9—12
stramineis
3
minus
Ad
39144
generally
coriaceous and
tepalis
stipitato, plus
v.
Cystolitha hypogena.
neata.
district.
longis,
mm
nullis. Flores feminei sessiles;
ovario sessili
ad carinam
persistentes,
appresse
1 —2
3,
costa
subgloboso, bracteis lateralibus nullis, apicalibus
(sicco),
servula
oblique, with
veins
petioli
brevibus, glandulis basalibus
1 —3
basalibus
lato
mm
12
Ramuli
elliptica sublanceolata
subsessilia, sparsim
bracteis
longo;
corpore
longae,
mm
lateral
longis appressis pallidibrunneis hirsuti. Ramuli
mm
—1.5
cm,
acuminate;
8.
spiraliter dispositis.
costis lateralibus utrinsecus
fuscibrunnea;
Leaf
348).
p.
Figure
—
copious.
F.
acuminata, basi subcuneata,
vix
attenuata
mm
Stipulae
12—24
(1969,
alta, foliis
m
bristles
shrub
Rheophytic
Corner
sp.
—io
parva
internal
(dried);
mm
—1.5
long.
mm
F.
3.
Figs
caducous.
of the lower side of the lamina
glabrous.
most
bracts
THE
long;
mm
New
more
on
fig,
crowded lateral
veins).
389C.
petioli
sp.
nov.
—
Figure
8.
rheophyticus, foliis spiraliter dispositis, habitu
costae
appressis
mm
lapidaria,
F.
Frutex
v.
subpatentibus strigosi. Ramuli
longae, lanceolatae persistentes.
mucronato
8 —12
mm
pallide fuseibrunnea;
intercostis
majoribus
ut
videtur terminaliformi. Ramuli
(ad paginam laminae inferam) stipulaeque pilis
—12,
vix
longo
costis
Lamina
acuminata,
lateralibus
elevatis, sparsim
subelongatis,
3—4
glandulis
mm
9—15
basi
appresse
basalibus
X
4
6
—
subeuneata,
utrinsecus
1 —3
mm
longis subfulvis
crassi, fuseibrunnei. Stipulae
7 —9,
puberulis;
obscuris;
cm,
obliquis,
apice
subcoriacea,
sicca
anguste
integra
14 —20
obovata,
subtus
valde elevatis;
costis basalibus utrinsecus
petiolo
8—15
mm.
1 —3,
Receptacula
BLUMEA
406
Figure
8.
Ficus
servula
(a)
and
F.
VOL.
XVIII, No.
lapidaria (b). Twigs,
x
2,
1970
½; fig,
x
2.
(NGF
39144
and
39036).
E.
axillaria,
J.
H.
solitaria,
subsessilia,
rubribrunnea (viva);
pedunculo
?
subglabris, persistentibus;
bracteis
lateralibus
longis,
albidis
sessiles;
tepalis
New
Corner:
pilis
—I
corpore
nullis,
—x
ligulato-spathulatis,
3,
celeribus,
rubris,
alt.
1500 m
This is close
with
to
fine
close,
F.
Hypogenae,
it would be
ficially
9,
the
makes
to
interesting
a
know if
novahibernica,
is much
remarkable
lepicarpa
F.
—2
mm
Flores feminei
sessile
v.
stipitatum
in fluminibus
ad
via Aiewa
parvis
Azeki,
lateral veins
they
parallel
of
with
copious
in
ser.
Amphigenae;
ser.
Dried material resembles
super-
Malesia.
western
subser. Congestae
(sect. Sycocarpus
oblique
more
rheophyte
a
F arbuscula in
with
together.
grew
are
provided
F. subcuneata. As
(sect. Sycocarpus)
sp. nov.
and
larger, villous,
Corner).
—
Figure
10.
Arbor
—8
alta, cauliflora
m
nodis
ccrvino-cinnamomei,
Lamina i8—30
data, integra
subtus
x
13 —20
cm,
axillis
videtur sine
glandulis; petiolo
sparsim
vivo
subnullo);
paulo
elevatis
mm
subanguloso,
ovario
v.
papillato-tunicatis.
46121
crassi,
caducae.
basi
7—17
saepe
2(v. 3), brevibus,
ad ramulos —3
caducis; pedicello
cor-
6—8(—9),
mm
1 —2 mm
X
1
cm
ut
efoliatos
longo;
bracteis
in
longo (?
statu
lato (sicco), turbinato, costis 6—9 longitudinalibus
lateralibus
nullis, apicalibus
4
—6
mm
longis
erectis
albidis numerosis; cellulis scleroticis nullis. Flores
breviter pedicellati; perianthio integro ad basim stipitis ovarii sub-
hilo
In
minus
silva,
stipitato; stylo setuloso.
subasperata.
prominenti,
Hibernia,
Nova
Cystolitha
Semina
I
Namatanai District,
Danfu.
longa,
mm
setis
hypogena;
internis
Typus:
NGF
(CGE).
This species
It differs in
in the
seems
most
nearly
related with F.
more
prominent apical bracts,
it is
not
F.
and F.
reoides,
seems
to
lead
novahibernica
nota.
and
to
may
In leaf form it
F.
tanypoda
and
the
stoloniferous.
longibracteata of the Solomons,
fig-orifice
tanypoda
Corner of the Solomon Islands.
being glabrous (except for the internal bristles),
figs; perhaps, also,
hand,
mm
subdichotomis etiam
versus
evoluta; pedunculo
mox
4—6
longae,
longo acuminata,
costis basalibus utrinsecus
longo. Receptacula
longis,
mm
rubribrunneo, plus
carinata,
compressa
bracteis
—0.4
mm
costis lateralibus utrinsecus
marginem
extensis
17 —20 mm
corpore
feminei subsessiles
cm
longis (ut videtur),
recurvatis; setis internis
cupulato;
5-—14
internodis haud
ramosos
3, 2 —3
5 —10,
22—30
mm
—15
brunnea;
elevatis;
vix
Ramuli
dispositis.
Stipulae
ovato-cordata, apice
glandularis,
saepe
intercostis
spiraliter
praeditis.
submembranacea levis, sicca
elevatis,
basalibus
foliis
glabra;
glandula
glandularis;
F.
fig
mainly the features of
are
lapidaria
the riparian
F.
437A.
These
internis
nullis.
lapides
Subdistrict,
Menyama
F. servula but the leaf is acuminate, the
intercostals,
internal bristles.
Inter
longis,
glabrescentia,
?
39036 (CGE).
NGF
Typus:
rubrum
ovarium
villosa,
mm
1.5 —3.5
setis
subumbonatis;
hypogena.
Guinea, Morobe District,
Nova
brunneolis
3,
pariete crassiusculo
in
superantibus; stylo valde laterali.
subundulatis
407
lato (sicco), subgloboso,
numerosis
cellulis scleroticis
copiosis;
(Moraceae)
bracteis basalibus
mm
—30
apicalibus
Ficus
and
longis
mm
longo;
mm
23
Streblus
species of
which
shorter basal
The
in the absence of lateral bracts,
bracts, and, possibly,
prominent apical
is also
bracts
the smaller
connect
it with
glabrous, and this general form of
the
that of subsect. Auriculisperma of the Solomons. On the other
relate westwards with
parallels
F.
emphasizes
their subseries in this subpachycaul
(elongate basal veins, persistent
cassidyana,
their
the
pachycaul
F. praestans, F.
significance
as
habit. The parallel in
basal
bracts).
ancestry
of F.
neobrittanica,
congesta
F. baccau-
local, relict endemics uniting
sect.
Neomorphe
is
F.
nodosa
408
BLUMEA
VOL.
Figure
XVIII, No.
9.
Ficus
2,
1970
novahibernica.
Twig,
x
½.
(NGF 46121).
E.
J.
H.
441A. F. cryptosyce,
Frutex
—1£
Corner:
sp. nov.
altus, sparsim
m
petioli receptacula
costaeque
5 —10
mm
Streblus
latice
ramosus,
cm,
v.
Integra
anguste
apicem
(ad paginam
longis,
laminae
inferam) pilis
superis
10—20
—10
mm
mm.
versus
vix
2,
elongatis, glandulis basalibus
I—2.5
mm
puberulis;
dein
longo; bracteis
corpore
7 —9
prominentibus, rugis
in
v.
pariete sparsis.
petiolo
2;
—18
5
glomerata, stipulis condita,
5
Flores
basalibus
mm
3,
2
—3
Figure
46121).
20—35
pilosae.
10.
15392
fusci-
longi
longae,
mm
Lamina
basi anguste
ÏO —23
X
subcordata,
costis latera-
costis basalibus utrinsecus
crassiusculo.
Receptacula
aurantibrunnea;
pedunculo
longo,
longis,
ovato-lanceolatis,
lateralibus
nullis,
appresse
apicalibus
feminei sessiles
v.
breviter
pedicellati; perianthio
v.
5
sub-
breviter
cupulato
stipitato; stylo sparsim setuloso. Semina
longa, subcarinata hilo prominenti subtuberculata. Cystolitha hypogena.
BW
II.
decurrentibus leniter elevatis; setis internis nullis; cellulis scleroticis
secondaria frequens; Nova Guinea, Vogelkop,
Typus:
mm
lato (sicco), bracteis
subinfund buliformi; ovario fuscirubro, sessili
0.8 mm
3 —8;
maturitate
Figure
crassi, internodis
fuscibrunnea;
sicca
mm
mm
Stipulae
longo acuminata,
subserrulata, subcoriacea levis,
—
mm
—0.5
—6
4
apprcsse
libus utrinsecus 6—8, obliquis, subtus elevatis; intercostis
axillaria binata
409
spiraliter dispositis. Ramuli,
glabrescentes. Ramuli
mox
obovata, apice
versus
(Moraceae)
Ficus
foliis
exiguo;
binac liberac late lanccolatae persistentes, basim
3.5 —8.5
and
(sect. Sycocarpus subser. Congestae Corner).
brunneis appressis rigidis hirsutae,
inferis brevibus
specks of
New
Lake
Ajamaru,
Tanah
Merah,
In silva
220
m
alt.
(L).
Ficus novahibernica.
a.
Figs,
x
I; b.
l.s. orifice
of
fig,
x
5;
c.
flowers
and
seeds,
x
10.
(NGF
BLUMEA
410
Figure
II.
and lower
Ficus
(3)
part
cryptosyce.
of the
a.
Leaves
plant,
X
½;
VOL.
from the
b.
figs,
XVIII,
upper
x
2;
(1),
c.
No.
middle
flowers
2,
1970
(2; stipules
and
d.
cut
seeds,
away
X
10.
to
show
(BW
the
figs),
15392).
E.
is close
This
There
F.
to
H.
Corner:
F. cryptosyce
seems to
axillary
shorter hairs
seed
and
Streblus
differ in the smaller figs which build
inflorescences concealed
Then the leaves of F. cryptosyce
(1
2 mm
—
the
(Corner 1967).
are
smaller,
be
only
may
subser.
to
by
Ficus
(Moraceae)
may
up
stipules.
I have
be
no
and both the
regional
Congestae they
be
a
as
a
small form.
living plant.
into considerable masses from
This is the habit which I have
record of it in F. multistipularis.
cupule
with shorter
petioles,
of the female flower and
differences of
may
411
merely
mostly entire,
narrower,
multistipularis),
in F.
Such
shorter.
are
I refer both
Though
species of
multistipularis of the Philippines and
illustrated for F. theophrastoides
the
New
few collections of F. multistipularis and little is known about it
are
the short
J.
a
widespread
derivatives of subser.
LITERATURE
CORNER,
E.
J.
H.
1962.
1965.
Check-list
1967.
Ficus
1969.
Ficus
HUTCHINSON, J.
in
the
sect.
1967.
The
classification
of Ficus
in Asia
Solomon
Islands.
Adenosperma.
The
genera
of
of
and
Phil.
Moraceae.
Australasia.
Phil.
Trans.
Trans.
Roy.
flowering plants
Gdns'
Bull., Singapore
Gdns'
Bull., Singapore
Roy.
Soc.
Soc.
B
B
256:
II. Clarendon
253:
19:
21:
23 —159.
319—355.
Press,
Oxford.
187—252.
1
—186.
species.
Axillares.