Download (Sapindaceae) Originally, my Soon, however, it considerably, finally

BLUMEA
28
(1983)
Notes
271-289
on
the extra Australian species
of
Dodonaea
(Sapindaceae)
P.W. Leenhouts
Rijkshcrbarium, Leiden,
The Netherlands
Summary
Australia,
Outside
D.
species,
restricted
elaeagnoides,
of
part
but
clear
elaeagnoides,
give
the
a
five
are
and
historical
are
and
of
were
All
for
a
the
of these
The last
complete
ber of
carya
of the
Small,
to
that
division
is
state
in
complex
used
by
Chapter
Pflanzenr. 98,
in that revision,
worldwide in the
main authors
with several
on
the
Originally,
Tropics
amounts
and
to
to
Radlkofer
varieties and
only
it
species.
to
name.
D.
In
and it is tried
III notes
given by
1933:
are
given
on
outside Australia
Radlkofer in his
1350—1404).
54, and
again
are
a
Subtropics.
into
a
a
further
and
one,
The
D.
species
to
num-
micro-
were re-
Madagascar,
Hawaiian species, and D. viscoWith Radlkofer the latter
was
number of forms and subforms.
E.E. Sherff
forms,
The Australian
intention was
to
became clear that
the
Madagascar
on
on
D.
viscosa,
the Hawaiian
species
this
enriching
and H.
species,
will be treated in
a
forth-
Ms. J.G. West, Canberra.
broaden the
liance all around
some
after
more
American
my
Soon, however,
part
Chapter
species
unknown to Radlkofer. All but four of these 55
was
coming publication by
least
three
to
D.
small
a
angustifolia,
mentioned
Australia, the exceptions being,D. madagascariensis, endemic
subdivided into three varieties and these
at
last
D.
after Radlkofer
revision of the genus Dodonaea is that
occurring
necessary
endemic
to
Introduction
microcarya, described from Florida,.D. stenoptera,
Lippold
into
In
or
the
Subtropics,
restricted
viz.
coastal
pantropical
and
IV.
sa,
The
polyandra,
under
a
Tropics
species three,
of confusion.
names
viscosa,
the
madagascariensis,
combined
of that
given
D.
Antilles,
and D.
time
species
names
the
D.
complex
viz. D.
accepted,
of these five
long
Sapindaceae (Engl.
species, accepted
stricted
of
Out
for the final
further
I.
monograph
part
Queensland,
for
given
are
species occurring throughout
to Australia.
explanation
discussion
a
and
viscosa
accepted species.
rejected;
inland
Florida
to
Guinea
D.
of Dodonaea
species
an
connections
II arguments
Chapter
to
New
Papua
with
five
angustifolia,
scope
revise Dodonaea for the Flora Malesiana
at
least for the D.
of my work
world. This necessity
attention
to
all the few
considerably,
made it
Ms. West's revision of the Australian species.
finally following
this al-
attractive and worthwhile
species occurring
area.
viscosa alliance it would be
outside Australia
as
to
pay
a counter-
BLUMEA
272
-
VOL.
28,
No.
2,
1983
II. THE DODONAEA VISCOSA COMPLEX
A. Radlkofer and after
far the
By
differences in leaf
the
gave
often
in Radlkofer's revision of Dodonaea is D.
complex species
most
variable and
extremely
and in
shape
mentioned by Radlkofer,
Tropics
and size of the fruits
shape
are
and
On
only, Radlkofer (1933)
(typical
forms and
necessary):
taxa
viscosa,
Subtropics.
added in brackets when
infraspecific
subdivision into
following
not
worldwide in the
occurring nearly
subforms,
Dodonaea viscosa
vulgaris (
var.
f.
viscosa)
var.
=
repanda
=
f-
(f. angustifolia)
f. schiedeana
(subf. schiedeana)
subf.
angustifolia
var.
viscosa)
waitziana
f.
thunbergiana
f.
microcarpa
spatulata
var.
f. burmanniana
(f. spatulata)
(subf. burmanniana)
(subf. spatulata)
subf. excisa
subf.
subf. laurina
Most of these
On
the
same
that
of these
many
After
to
the
two
Finally,
synonymy
few
a
D.
Sherff (Field
(for
an
weakness
eriocarpa
1951:
placed
D.
as
a
course,
D.
rather
on
distributed.
widely
in the
occur
makes the
already
impression
charac-
unimportant
mainly
on
is
Hawaii,
eriocarpa
of these
forms
it
a
admittedly arbitrary,
viscosa.
Radlkofer!),
One
of these
became
subdivided into
was
John,
D.
some
as
a
D.
note
viscosa
25—30 varieties
complex
Sherff
by
newly
complicated
List Flow. PI. Hawaiian 1., 1973:
223).
and the D.
(Amer.
J. Bot.
38,
coherent group of Hawaiian forms
eriocarpa notwithstanding
belong
Radl-
eriocarpa.
oxyphylla):
rather
varieties
placed by
taxa, viz. f. elaeagnoides and subf.
demonstrated in
f.
names
happened
by
between the
even
some
1947: 269
more
later authors. This
subforma with
distinction
clearly
revived
described around
were
several
by adding
Furthermore,
were
infraspecific
whole under D.
while
ones.
viscosa
eriocarpa (a
of the
59, sub
are
Mus. Nat. Hist. Bot. Ser. 23,
this system
enumeration see H. St.
complex
some
of D.
species
new
viscosa itself as,
and forms
and
phenetically,
complicated
of Radlkofer's
accepted species,
ters
further
forms, partly again widespread
also
The
defined
are
local endemics but
no
of these varieties and forms
may
different habitat. This
a
Radlkofer, especially
kofer in
as
in
are
eriocarpa
elaeagnoides
genetically.
enriched and
317)
taxa
taxa
several
to
some
often
though
ters, rather than
and
infraspecific
other hand,
area,
f.
under D.
has seemed the
the fact that
viscosa
as
as
defined
only practical
one
to
by
to
is
the charachim. 'This
employ.'
he
argumented.
The
only regional
pold (Wiss.
revision of
some
importance
Beitr. Friedr. Schiller Univ.
the American
species.
zonica, D. bialata, D.
He
divided the D.
elaeagnoides,
D.
since Radlkofer is that
Jena, Beitr. Phytotax.
viscosa
complex
linearifolia,
and D.
6,
1978:
by
H.
Lip-
79—126)
on
into five species, viz. D. ariviscosa.
P. W.
B.
My
In
Flora
Radlkofer's revision
A
three
stricted
small
in
a
widespread
viscosa
apparently
forms,
species
f.
or
followed
by
One
recognized.
of the Western Dist.,
one
1945:
var.
minor and
de-
212)
var.
D.
was
this
angustifolia,
re-
.
one
was
D.
representing
form, mainly
a montane
was
there
Secondly,
sometimes andromonoecious and bears lanceolate
Radlkofer's D.
mainly
comparison,
polyandra,
viscosa f. burmanniana, sometimes his
mainly depending
on
forms,
two
and
eriocarpa belong,
the relative width of the
hairy
a
glabrous
a
or
one
on
Java,
nearly glabrous
in New Guinea.
The
morphological
well
as
the
as
differences between the second
ecological
the third entity mentioned above did arise doubt
of Radlkofer and of Sherff. The
tems
in
to
known
32,
careful mutual
a
of these
New Guinea.
Papua
system. The third
or
which Sherff's varieties of D.
one
of
Guinea, described after
coastal form with obovate leaves and bisexual flowers,
represents
was
the other
coastal,
one
varieties, viz.
two
leaves. This third group could further be divided into
to
viscosa, which
New
from
According
matter.
Sherff (Amer. J. Bot.
eriocarpa
material,
entities could be
f. repanda of Radlkofer's
schiedeana
rare
the revision
above,
easy
Java.
on
Malesian
part
an
and variable D.
furthermore,
Hawaiian D.
be
to
different
slightly
two
E. Malesia, which is dioecious
leaves; this
f.
273
Dodonaea
species of
taxonomy sketched
seemed
area
widespread
a
out;
came
of the
learned that
a
extra-Australian
complicated
an
mainly
occurring
as
sorting
to
of
area
polyandra,
scribed under the
waitziana,
on
Malesiana
could expect
that
in
montane; D.
the
with
the
the literature I
consist
Notes
approach
own
comparison
Dodonaea for
to
Leenhouts:
hairiness appears
be divided
hand, however,
form,
were
clearly
is
system
moreover
that
it
flower. Therefore, I decided
distribution in order
to
form, which apart from the variation
montane
with Sherff
even
get
to
based
over
two
the D.
insight
on
in the
main
through
of these
variability
for
shape,
this
with
differences in
all
complex
other
montane
cause
variations in leaf
made of the
was
viscosa
should
On the
species.
viscosa. The
mainly
mention
no
study
better
a
vulgaris of D.
var.
was
and
of the sys-
differententities, the coastal form and part of the
pubescence;
on
correctness
well-circumscribed taxon, in Radlkofer's
system
a
three varieties,
both included in the
unsatisfactory
Sherff
two
be
to
all
over
regarding
the
two
its
the
area
of
forms and
in the constancy of the differences found in Malesia.
The
Only
they
coastal
in the
were
tionally
always
The
form
appeared
Caribbean
a
monoecious,
few
or
to
low altitudes
inland form
nearly pantropical
had
along
appeared
or
just
behind the
to occur
variation in leaf shape and size,
populations
from Tahiti and E.
growing under rather arid
South
Africa,
and
clinal variation
or
the
not
and is
with
Malesia,
conditions
surprisingly
exclusively,
the flowers
were
Sto.
New
on
on
all
or
Domingo)
partly
nearly completely
func-
it occurred
sea coast.
Zealand).
the
one
It showed
hand the
a
lying
as
far
apart
relatively
as
flowers showed
dioecious
(Califor-
wide but continbroad-leaved
the other hand linear-leaved
in areas
uniform.
bisexual flowers;
all around the world between 35° NL
the southwestern U.S.A. The
from
or
exceptions (the Seychelles,
nia, Arizona) and 40° SL (the South 1. of
uous
no,
andromonoecious,
female. With very few
at
be
specimens
populations
southern Australia,
a
populations
rather continuous
from the
Pacific
274
Galapagos I.), Australia,
the
(except
andromonoecious
the
to
monoecious
or
predominantly
tane
or
submontane
the
their
here
even
forms
two
The
area.
in
the
Summarizing,
that
hold
not
seen
final
My
C.
which enable the easy
any
is that
allied
distinctly
separated
nearly throughout
in America,
completely
identification of
a
between the
hybrid
partly
due
each other and
meet
never
several
specimen
every
forms may be
two
they simply
nearly
though
are
two.
the coastal and the inland form should be ranked
The
species.
that for the inland form
Jacq.,
do
mon-
about 20° NL and SL.
at
is
D.
correct
as
for the coastal form is
name
L. f.
angustifolia
Before Radlkofer
argumented above,
As
conclusion that this
and
viscosa
is
question
the
how
species
it
second question,
was
the
any
The
name
first half of the
The
so.
20, pi.
He
on
first
the
to
Dodonaea.
so
—
use
the
very
a
name
(Gen. PI.,
that genus in his
reference
to
found the
1763:
In
complex.
91, being
342),
then
illegitimate
1737, while rejecting
341) proposed
diagnosis
of the
that
as
the
a
in
by
the
later
name
same name
flower, apparently
a
was
as
species
could be answered
only
from this
plants
already
both
were
in the
named
Plant. Amer. Gen., 1703:
this
in the synonymy of Ilex.
which
name
could
easily
escape attention: in the
script (accepted names)
Dodonaea in italics
sense
The
of Plumier
homonym
quite
accepted
points resulting
different
with
another
was
sense
based upon dried
reference
Adanson
of
and
(Fam.
PI.
a
I
2,
(1754).
Plumier,
He gave
material,
to
with
later author where
only
plant.
a
(rejected names)
of Dodonaea Miller
Dodonaea in the
for
A
part in the systema-
viscosa s.l.
questions
the number of the genus Ilex.
Dodonaea used
name
way
logical
species separate.
Plumier (Nov.
index) placed
Roman
be well
subchapter.
completely
Dodonaea was
followed
sense,
These
the
is whether and how far the
played
As D.
to
A first
is whether the division of that
question
two
inconspicuous
own
has
1978).
these
troubles around Dodonaea started
1737:
index he gave first Dodonaea in
Lippold,
one,
also
to
from D.
apart
as
appeared
of literature. The main
study
The
H.
first
polygamy
in this
typification.
18th century when
Linne
12).
did this in
PI.:
accepted by
flowers and
especially
influence
elaeagnoides
hardly anybody kept
will be mentioned and discussed in this
study
a.
that
of Radlkofer led
sense
species (actually three,
connected with the
extensive historical
fairly
a
already
of its
the third
genus,
type species
will have
and
viscosa in the
two
Caribbean D.
possible
difference between bisexual
the
included
was
distinctly
tics of Dodonaea,
of D.
study
complex
angustifolia
D.
but this
separable,
by
that could be
specimen
conclusion
different, though
viscosa
well
are
bisexual
rarely
not
nearly exclusively
of the material. Moreover, there
majority
a
small part
a
all around the world learned
complex
differences in distribution and habitat:
have
D.
viscosa
in Malesia
be
to
the lowland
the flowers won't
for
via the for
dioecious,
sometimes
The clear differentiation between the
(see Chapter III).
I
1983
bisexual condition in South Africa
exceptionally
of the D.
study
good
2.
Malesia and southern Asia
Tropics, reaching
distinguishable
additional characters
the
the
No.
28,
habitat, this form appeared
to
differences in
they
to
VOL.
andromonoecious,
condition in America. As
to
-
BLUMEA
a
as
Linne
fairly
to
(Gen.
extensive
older litera-
P.W. Leenhouts:
he referred
ture
he gave
144)
237)
1747:
gave
sis of 1737. In
based
PL:
viscosa L. The
differs in
54)
and is
absolutely
Linne
did
(Gard.
ler
inal
all.
at
the
Diet. ed. 4,
first
than
more
162).
same
diagnosis
donaea,
is
mainly
and differs
Dodonaea
name
viscosa.
Like
he refers
ferences in the
nosis from
(Select.
‘Mihi
(Mant. PI.)
(1763).
be
out
').
sum’
end,
Linne
The
of
no
a
diagnosis
109, sub D.
60)
b. The
as
This is
Miller
possible
angustifolia.
Syst. PL,
was
of
1760:
27, pi.
followed
however.
former,
a
synonym of Ptelea
short
(‘Ptelea foliis
comments
the dif-
on
a
rather extensive flower
is
Syst.
with
a
ed.
he
completely:
sur-
diag-
12
based
in the
typification
the
(p. 228),
Pteleae
original
sense
1754)
sense
of the
of Linne
viscosa
genus. Linne
described the
accepted here,
pistil
but far
named under Dodonaea, i.c. D.
exclusively
on a
into
reference
to
in 1771
original
Jacquin
appeared
plurium
Dodonaea
name
duplica-
Plumier's
upon
which
genere
slightly
the
Finally,
based
mainly
The division of D.
Diet. ed. 4,
species
on
Dodonaea in his
accepted
a
125.
(1767)
the confusion around the
the
original descrip-
viscosa solis staminibus
description
(‘Removende itaque
Dodonaea.
for
extensive
made of Dodonaea in another than
149)
viscosa in
first
19),
botanists,
own
was
nothing
1747. With Linne, the
as
extremely
mentioned is Dodonaea viscosa
(Gard.
in D.
The
still
viscosa) published
generally accepted
consequences
as
(p.
publication
typification
well
his
on
orig-
additions derived from Do-
Linne for the first time commented
his mind
under Ptelea
and the genus
is
not
Mil-
19)
1725:
212) hesitatingly
of the
an
several authors, and finishes with
by
No word
mention is
genus
only species
With this
173),
is
sense
with
(Jamaica 2,
29)
In 1754,
1760:
viscosa,
he gave in
of several
publications
of the flowers
PI. ed. 6:
changed
as
place
could have
341)
Hist.:
differre’);
of 1737
sense
the
D.
name
some
PI. ed. 2:
The last remark holds also for Linne,
sense.
1760:
of Ptelea viscosa in that genus CP.
position
videtor
(1753)
own
Syst. PL,
i.e.
Sloane
under the
diagnosis
(Sp.
(1737).
1763, Fabricius (Enum. ed. 2: 430, sub Dodonaea) and Jacquin
Amer.
tions. In 1764 (Gen.
aberrant
1762
floruit’.
non
1737! In
Stirp.
in
the
to
descriptions
remark:
prising
from the
completely
reappeared
PI.:
(Gen.
Linne's Hort. Cliff, and with
to
Gen. PI. ed. 3,
Ptelea,
PL ed. 5,
(Gen.
validly published by
was
Jacquin (Enum.
based upon Ptelea, but with
the first edition
in
simplicibus’);
Ludwig (Def.
synonym of
a
1737
Dodonaea in his
Dodonaea
palituro affinis
Dodonaea and
combining
Linne
in
given
as
of Dodonaea
under Dodonaea,
name
Aceri vel
to
year
reference
a
flower and fruit.
species
a
reference
a
In the
Linne in
His
publish
to
with
1754)
name
however,
year,
given by
one
original diagnosis
2,
diagno-
(Linne, Sp.
1753
regarded
is
now
from the
only
reference, and the
same
description including
the
tis
few additions
any
In
of Linne
sense
Sta-
Cliff.:
Gen. PI. ed.
based upon Linne's
references. In
new
to
(Hort.
apparently original description
a new,
some
and
pi. 23)
In 1738
18).
probably
of the genus Ptelea
diagnosis
a
different from his
give
not
mentioned
the
in
275
vegetative parts. Ludwig (Def.
of the flower,
and he added
Dodonaea
name
1737: 55,
1703:
Gen.,
1748, Linne (Fl. Zeyl.: 58) gave
the
118)
1754:
diagnosis
species of Dodonaea
(Thes. Zeylan.,
of the
description
short
a
extra-Australian
Amer.
Plant.
(Nov.
fruiting material,
on
Ptelea
short
a
on
Burmann
Carpinus
to
Plumier
phylodendron
Notes
came to
three
being
species
1737:
3-merous.
more common
viscosa
Sloane
an
(1737).
(Gen. PI.,
as
to
consen-
in D.
Jacq. (Enum.
(Jamaica 2,
1725:
276
BLUMEA
Sloane included
27). Apparently,
the former
mainly
latter
a
of the fruit
mixture of the characters of the
inland
The
population).
his herbarium. According
the
Harbour;
viscosa, the other one,
proposed
this is
The
c.
sulting
long
this
as
the
acters.
sed in
in
for this
from
a
description
bisexual
whether
Miller
1756:
and
case
(Gard.
Linne
In
angustifolia.
the
two
that mention
part
G. Forster
only,
This
now
1737:
all
(Fl.
may
have
Sloane
341)
were
ones or
were
was
plate
finally
re-
polygamous.
or
value of it
understood,
Finally,
overrated.
less
or
built up
when
the fact
notwithstanding
more
already
understood
mainly
on
by
leaf char-
will be mentioned and discus-
and 207,
In
well
were
Dodonaea),
Dodonaea),
(Suppl.:
in the
218)
diagnosis
(Civ.
Fabricius
Amer. Hist.,
237,
the
1786:
to
based
viscosa
27).
was
He
insulis
upon D.
only
D.
failing
knowledge
angustifolia
this
(Gen. PI.,
1763:
next
1778:
D.
species,
leaf
be-
shape
the first time
was
but
this could
step
was
not
taken
by
viscosa, described the leaves
hermaphrodita.’.
character of D.
of that time
occurs
ed. 2,
second
a
form from the
angustifolia (the
PI. ed. 2,
Dodonaea),
109, sub Dodonaea
dioica. In Nova Zeelandia
the confusion: the
broad leaves and could with the
D. viscosa; in New Zealand
unknown. The
mentionedD.
(Gen.
sub
difference in
hand, however,
being
Nat. Hist. Jamaica,
and Reichard
described
exten-
of
descriptions,
(Enum.
1763:
palituro
rather
impression
Linne
1747:
P. Browne
a
flower
polygamy (' (‘Fructificatio polygama.’),
In Soecietatis
only
published by
Dodonaea),
is stressed. On the other
Ins. Austr.,
was
as
give
to
Comparable
Gen. PI. ed. 2,
1771: 149, sub
as
the first
the flower makes the
viscosa.
(Def.
1782 Linne f.
made of
added
D.
sub
1725: 27, sub Aceri vel
2,
apparently
original,
Ludwig
the situation in D.
given, but it
typical
number
and with the
bisexual
systematic
leaves
(Jamaica
was
Jacquin (Select. Stirp.
name
species
'
specimen
the insufficient under-
probability
character
described,
1754: sub
(Mant. PI.,
was
as
of the
of this
representing
the older
the
As
description
important publications
by
note
Triopteris,
but added:
relatively
as
more
Diet. ed. 4,
191, sub
shown
58-59)
cited under
v.97, they
of the systematic confusion
to
influenced the system
accordingly
194, sub Dodonaea).
a
cause
was
and size
shape
159, sub Dodonaea),
viscosa),
play
complex
of the flower. As
repeating
is
order.
short
430, sub Dodonaea),
tween
The main
—
well known, let alone the
not
(Gen. PI.,
Linne
of the
sense
1926:
but
to an
viscosa and both with
angustifolia.
var.
points the
anything
be
Jamaica 5,
(Fl.
nr.
one,
angustifolia,
species
two
collections both named D.
The
some
viscosa in the
differentiate between the
Hills'.
D.
in
'Red Hills' may refer
D.
depicts
as
hardly
can
the flower conditions became known, and
chronological
Apart
1742:
locality
the
species;
well
as
harbour'),
4-merous
to
v.99, under the
systematic importance
The
affinis),
sive
was
botanists, it hardly
some
'Old
lecto-type.
differences in
variation
that
viscosa
of the flower conditions, whether the flowers
especially
the
D.
locality
2-
1983
with the main part of the
conditions.
flower
two
in the Dodonaea viscosa
standing
As
as
the
not
Red
nr.
is best in accordance
v.97
two
2,
Fawcett and Rendle
to
Sloane herbarium includes
the localities 'Old
D.
text
being
as
No.
28,
plate (nr. 162) clearly
present author. That Sloane did
by
in his
of the leaf;
(description
(the description
VOL.
-
and it
only
seems
viscosa
Society
was
I. has
be identified
very
improbable
P. W.
that Forster
1791:
Bot.,
would have
150)
the first
was
one
hand gave
tree
(7n
(Icon. 4,
in his
but
D.
1797:
known,
Cycl.,
His
incorrectly
listed
gamous D.
description
actually
and
he added
plate clearly
as
species,
described
for D.
same
described
DeCandolle
Moench,
(Mem.
D.
representing
Soc.
Geneve
as
merated
17
he
species;
cordingly
defined his
Sprengel,
in
cluded
the
became
a
only
true
D.
the
described the
species primarily
9th
edition of Linne's
'
diagnosis
genus
genus
genus
on
character,
‘Flores
(Rumphia
188)
ferent conditions of the flowers
sexualibus, monoicis
v.
dioicis’)
Genera
flower characters
as
useless
tant
sometimes
13, 3:
though
d.
on
abnormally
misunderstanding
639)
he
he did
The growth
Dodonaea had
make
of
(1830:
time
condition
viscosa.
Moreover, before
disposal.
next
complex.
This
means
1782,
that it
—
this
mainly
was
to
divided
him,
and
ac-
in-
it
v.
abortu
‘Bl.
356) regarded
diöcisch
the last
(Mart.
viscosa
impor-
Fl. Bras.
one,
complex.
publishing
and Asia,
mainly
angustifolia,
angustifolia,
(da-
more-
hermaphrodite
D.
difuni-
of his
description
was
either D.
was
described the
Antilles,
described D.
was
enu-
the fruits.
on
The first botanists
represented
when Linne f.
a
growing
the
Rox-
to
for that purpose, the
species:
to
poly-
appar-
318) simply
Pflanzenfam. 3, 5:
of it for the division of the
material from America,
at
445)
hermaphrodite)’ (Flowers dioecious,
the Dodonaea viscosa
only
the
to
the flower conditions
correctly
But
the
to
DeCandolle
hermaphroditis
....
un-
Rees
1814: 28;
polygamous,
or
species;
for the distinction of the
polygamous
a
(in
ample opportunity. Finally,
seemingly hermaphrodite).
use
Ceylon,
their
616)
of the flower conditions in Dodonaea: in 1900
recognized
not
still
what different
unknown
and included this character in the
neben anomaler Weise mitunter scheinbar
over
1822:
angustifolia
In this way
who for the first
possible (' (‘floribus
contrast
1,
secondarily
Plantarum
species. But Radlkofer still in 1895 (E. & P., Nat.
the
D.
leaf characters,
polygami’.
and the
dioica ascribed
1824:
(1,
unfit for the distinction of
1847:
3,
as
dioecious
as
he
angustifolia,
plant
but added
Furthermore,
angustifolia,
wide variation of the leaves and the fruits offered
Blume
D.
a
Wendl.
triquetra
as
mention them
D.
with Linne f.
dioeca in
Phys.
doubtful. In his influential Prodromus
regarded
in
species;
some
not
to
Roxburgh (Hort. Beng.,
species and described
Roxburgh's
DeCandolle into
refer
Cavanilles. Smith
and
dioecious D.
a
same
viscosa. Cavanilles
being dioecious,
some as
the
on
the
on
the confusion
to
viscosa,
authors from different parts of the world had described
by
growing
variation, and named his plant
a mere
viscosa.
treated
was
358)
1794:
of D.
synonym
female flow-
or
the differences in leaf shape between his
by Forster,
256)
angustifolia.
but
illegitimate
filled in
the
1832:
2, 2,
misunderstood
burgh
hand, however,
(Obs.
from Jamaica. He
viscosa
(Methodus,
flower condition of D.
the
seven
Moench
Swartz
area).
angustifolia
of male and either bisexual
Sloane he considered this
by suggesting
Fl. Ind. ed.
on
from that
of D.
male and female flowers but did
pi. 327) depicted
Accordingly,
1809)
flowers
of male and female flowers
an
277
species ofDodonaea
from different shrubs. D.
were
lucida,
he commented
was
confusion
ently
D.
original description.
viscosa.
they
On the other
stirpe’).
depicted by
one
that
without flower characters.
species only,
though
extra-Australian
hermaphrodite
original descriptions
eadem
one
had
on
good original description
a
indicating
and
briefly
gave
gave
Notes
give separate descriptions
to
ers, moreover
very
Leenhouts:
or
D.
Dodonaea
BLUMEA
278
considered
was
a
mixture of the
D.
presenting
from
1)
Ceylon;
from
consist
to
given by
scriptions
of
species, hardly
angustifolia
Burmann's
Acer minus
Plukenet's Arbuscula viscosa
the
58),
second
one
from
Ptelea
America;
Jacquin's
ca
reus
species
two
Amb.
(Herb.
with
Starting
tween
Lamarck
ed.
(Rees Cycl.,
B.
D.
The
(Fl.
several
authors into 5
added
as
Sprengel
a
as
it led
growing
& K.,
1824:
but
ed.
some
was
lian,
Sp. 5,
1821:
variable
qu.
authors
lito-
be-
distinguished
&
150), Murray
2, 1,
to
a
are
Persoon
and Smith
343),
1799:
ed.);in
D.
leaf characters
unknown
to
wrong
naming,
these three
the
him).
(the
This
description
the
cases
angustifolia.
and G. Don
242)
to
the
D.
viscosa
as
extra-Australian,
complex
(the
was
same
'The
as
form of the
as
forms,
133
laid
well
as
true
D.
system
Soc.
(Gen.
as
Phys.
such
angustifolia
by
was
followed
was
Hist.
1831:
1,
of some local D.
that of the leaves'
ed.).
He
Bentham
the
are
the fact
wings
species,
by
673);
angustifolia
subdivided into
by
of the
(p. 472)).
strength
(Fl.
angustifolia).
Austral.
of much
1,
newly
three named varieties,
nearly exclusively
1863:
As
a
(in H.,
viscosa
The basis
475),
species,
how-
Austra-
collected material.
based upon leaf and
used for the identification
that in the introduction
capsule,
Knuth
divided what he called D.
the addition of several named
divided into
characters
variable
big
taken for the first time
all three represent D.
by
by
on
one
was
qu.
(actually,
was
species, notwithstanding
marked:
133 of the
on
other botanists
final system
enlarged
fruit characters
of his
from
palituro affinis (Jamai-
he divided the material identified
616)
Dodonaea viscosa
Nov. Gen.
but also
Finally,
PL ed. 4,
clearly represented
1825:
number of varieties and
of Radlkofer's
He
from
109)
started with DeCandolle: in 1822 (Mem.
mainly
16, 2,
into three unnamed varieties
ever.
1763:
species.
The other way,
B.
1,
species,
one,
(Syst. Veg.
moreover,
forms
sixth
probably
fig. 2)
angustifolia. Examples
1791:
(Obs. Bot.,
1821:
Sp. 5,
angustifolia
Prodr.
445;
1:
Swartz
used for what
of D.
247
Zeyl.,
P. Browne's
Austr., 1786: 27), Cavanilles (Icon. 4, 1797: 327), and Knuth
Ins.
was
splitting
several
218)
narrow-leaved D.
a
Willdenow (Sp.
386),
Ceylon;
149, 228) from America.
1771:
This unreliable character led other authors
Nov. Gen.
& K.,
viscosa
Geneve
1797:
15,
18)
from
144; Fl.
probably Rumphius's Caryophyllaster
1782:
and
8)
fig.
1703:
from Jamaica;
Hist.,
141
Ceylon;
Ceylon;
245, pi.
Amer.
Stirp.
pi.
from the Moluccas.
(Suppl. PL,
292),
fig. 1)
re-
1689:
from
32)
1738:
from
Sloane's Aceri vel
up in
pi. 50)
viscosa
1786:
2,
1809).
e.g. G. Forster
name
Linne f.
(Encycl.
(Syst. Veg.
(in H.,
1743: 110,
4,
1692:
pi. 23)
55,
1760:
(Mant. PL,
some to
clearly
names
(Cat.,
1717:
Cliff.,
one
18
Amer.,
(Select.
from Jamaica and
broad-leaved D.
a
mixed
apparently
are
of
angustis
1736:
191, pi.
simplicibus (Plant.
fig. 3)
1725: 27,
2,
lauri
foliis
America, the third
Dodonaea viscosa
angustifolia,
Zeylanicus (Prodr. 2,
1737: 341; Hort.
Linne's Dodonaea viscosa
America; and
The
foliis
that the de-
appeared
D.
Examples
(Mus. Zeylan.,
(Thes. Zeylan.,
(Gen. PI.,
Triopteris (Civ. Nat. Hist. Jamaica, 1756:
Plumier's
it
to
Elaeagnifoliis (Phytogr.,
Waerellaghas
viscosa
Carpinus forte
Linne's Dodonaea
refer
viscosa !
America; Plumier's Staphylodendron
probably
1983
2,
clearly
D.
to
any
Breyne's
are:
No.
species only. Surprisingly,
one
also from America; Hermann's
1748:
28,
of the older authors
most
two
VOL.
-
to
the
genus he
which has been much relied
for the reduction of several
on,
is
names to
re-
as
D.
P.W. Lecnhouts:
viscosa he
was
latter gave
no
by
followed
at
off and
1900
in
new
regarded
was
and
D. angustifolia.
The D.
the
viscosa
D.
more
with
complex.
viscosa
ferences
known
a
more
D.
two
Java, but they
and D.
in the D.
species
beach form and
been
paid
a
in
and validated
to
was
the
(see
by
considered
are
an
3a.
when
bisexual.
spicuous
Tropics,
b.
Scar
least
annular,
morphological
In
at
least
ecological
dif-
Java, he distinD.
a montane
ferrea (not
ecological,
represent
Bakh.
Fl.
f.,
mentioned but
not
remained well
mere
Java
forms of
one
2, 1965: 141,
named).
THE ACCEPTED SPECIES
TO
THE SPECIES
3. D.
12—15;
stamens
madagascariensis
9
flowers
without staminodes. New Guinea,
4. D.
stamens
5—9; 9 flowers with
dried rather slack,
of
stamen
scars;
papyraceous
under the fruit
sepals
the
fruits
or
to
3
Flowers
thin-pergamentaceous.
mostly strongly
mostly 2-merous,
lobed around
strawcoloured
or
the
stiff, pergamentaceous
to
often dioecious. Scar of
slightly
lobed,
scars
of
chartaceous. Flowers
sepals
under the fruit
staminodes
often
partly 3-merous, sometimes 4-merous, especially
when
mature
viscosa
mostly
usually
inconspicuous;
the
con-
brownish.
5. D.
partly unisexual,
sometimes
polyandra
without staminodes
coastal
Leaves when dried rather
at
to
distinctly
2
5;
Sepals mostly 4;
Leaves
much.
indirect reference. The dif-
Queensland
b.
of
of the
Tropics.
Leaves pinnate. Madagascar
mostly
were
thanks
latter is
role
aware
and
to
e.g. Backer &
mountain form
the
and
morphological
always
viscosa
KEY
Sepals
the ad-
change
worth mentioning, however, that
(never described)
taxa, both
III.
simple
(in
paper.
complex,
viscosa
not
does
angustifolia complex and,
angustifolia
montana
were
and variable D.
Leaves
1933
species.
viscosa, actually
a
in
complicated by
its varieties and forms, the
It is
coastal D. littoralis
between these
on
widespread
where
a
viscosa
later renamed D.
described),
2a.
all
attention has
chapter
between D.
between
guished
b.
three
recognize
and D.
allied
Finally
IIA of the present
the other hand parts of it
on
author, i.e. Junghuhn (Java ed. 2, 1, 1853: 296),
ferences
la.
closely
but the
697),
than the former ever was!
this
characters in the D.
one
to
279
further elaborated
was
13, 3: 639),
Subchapter
whereas
replaced by
now
eriocarpa
unmanageable
Throughout
is
Bras.
hand made still
one
elaeagnoides
complex
addition of D.
Fl.
forms,
distinct, though
as
Bentham's system
Mart.,
the
on
varieties
species of Dodonaea
J.D. Hook., Fl. Br. India 1, 1875:
(in
complex.
(in
The present author's decision
viz.
extra-Australian
on
in the form given in
1363)
that system
dition of several
split
first
Pflanzenr. 98:
Afterwards,
Hiern
by
subdivision of the
Radlkofer,
Engl.,
Notes
wings tinged
fruits
reddish
4
BLUMHA
280
Staminodes
4a.
incision
ly
fruits
conspicuously
and
incision
Fruit
Dodonaea
L.
angustifolia
tifolia Benth., Fl.
J. Bot. 32
Amer.
1983
2,
wings
adnate
not
fruit
the
to
style, according-
1
PI.
the
the
to
style
fruits
body;
West Indies
218.
(1782)
D.
—
—
viscosa
D.
angustifolia
and
accordingly
mostly
viscosa
f.
Jacq.
sparsely
D.
2. D.
not
con-
elaeagnoides
Jacq.
angus-
var.
angustifolia Sherff,
(LINN,
herb. Linne 495: 4 and5
Type:
—
adnate
wings
reaching
476.
(1863)
214.
(1945)
sometimes
glandular,
1.
not
f., Suppl.
Austral.
densely
not
spicuously glandular, glabrous. Florida,
1.
No.
inland
flowers. Fruit
top of the
on
often
though
Subtropics,
Staminodes absent in 9
the
28,
of the fruit;
top of the fruit between the wings reaching the body
on
hairy. Tropics
b.
flowers.
9
present in
VOL.
microfiche
seen).
Most of the
rejected
The
demarcation between D.
Therefore, and
variability
order
upon
more
(—12)
blackish than in D.
times
as
long
stiff pergamentaceous, alive
sides,
6
as
big
tent.
as
flowers
3-merous,
halfway
wings)
or
10°
on
at
halfway
Pistil
open
rocks,
NL and
were
a
11
to
vegetation
rarely
SL
to
not
low
(difficult
1—2.5
on
is
the
to
1.5
bisexual.
cm,
mm
or
hardly
mostly
forest
mostly
along rivulets,
1000
m,
in
reaching
of 25
out
(part
the
the
of the
anthers,
plants
are
in
fairly long persis-
2—3
mm
one
in 6
fourth
body
reddish
as
well
specimen
on a
many
Fruits: the
on
as
case
often
for
so,
densely
for certain,
espe-
female
to
some
so.
both
on
mostly narrowly
say
long, mostly
style
3—6.5
Sepals 4, exceptionally
lobed, sepals
to
some-
middle, pergamenta-
andromonoecious
were
fully
shrubby
etc., also
always
x
varnish. Flowers:
shining
wide; rather stiff,
above
the
indehiscent; in that
are
be
several 4-merous;
mm
12.5)
—
in
probably functionally
specimens
the apex,
steep slopes,
25°
notes
acuminate. Inflorescences
normally 2-merous, often
to
3.5—6
to
the latter
rounded,
tapering
poor),
are
appeared
bisexual
some
by
when young and the bark may
exceptionally pale brown, the body conspicuously
Habitat:
often
cleft
to
under the fruit annular
some
following diagnosis
and inflorescences
bases,
elliptic, 4—9(
dioecious
apparently
specimens
and 8
flowers.
sometimes
high, wings
to
scar
Anthers in
in bisexual
than
5;
acute
flowers, apparently
in d
as
the
is followed
slightly coriaceous, glands conspicuous
sometimes
be bisexual
to
actually monoecious),
some
were
if the herbarium
flowers considered
leaf
glandular
all branches covered with
flowering specimens
though
apex
broadly rounded,
hardly glandular,
cially
more
viscosa. Leaves
leaves often varnished;
to
diagnosis
wide, widest somewhat above
as
to
if
be
viscosa is weakest in America.
comparison,
high. Young parts,
ceous
mucronate
to
considered
are
its area.
m
Branchlets may be
slightly hairy.
chapter
next
and D.
mutual
a
American material. This
2—6
shrub,
or
the
angustifolia
facilitate
to
in other parts of
Treelet
be
in
based
exclusively
times
discussed in
names
angustifolia.
synonymous with D.
to more
7—14
mm
(especially
the
glandular, glabrous.
dry
soils
(sand, limestone),
secondary vegetation
etc.
Up
the lowland
20°
NL
at
about
and 25° SL.
Variation.
conditions
In the
seems
to
western
be
part of the
clinal, in
the
area
eastern
(America, Africa)
variation
the flower conditions
part
in
are
flower
nearly
P.W. Leenhouts:
pattern; partly
hold
for the
true
Locally,
the
taxa,
extra-Australian
it may
of variation
it
areas
though
depend
most
warranted
When
show
continuous
a
viscosa. Nearest
Out of 14
and
flowering specimens
bisexual
whereas
well
flowers,
only
one
developed
des, but
one
well
are
The further material from
Australia, and the Pacific
times monoecious
or
American D.
the
same
described from the
from
sterile
one
leaves,
vate
and the staminodes
Most
two
H.H.
van
apiculate
glabrous
are
The
which has
Celebes,
are
viscosa
der
dd),
varied
at
var.
flowers,
from rather
non-dehiscent.
and
apex,
I. agrees
developed.
some-
exception:
excellently
or
9
with
flowers
on
The other form
is known
to
me
only
This has rather broad obo-
(U).
resembles
indicate that the
possibly
notable
one
galapagensis,
1914
had d
one
for the stamino-
normally
and either d
rather well
Werff
main
strongly Degener
28355
I. received Do-
Galapagos
especially
the inflorescences
only
or
is the
exception
of the
pistils,
thinly, shortly,
are
form from Java, the
and
infructescences
of the
accordingly
densely
is
fruits,
and
Lesser Sunda I., and
the leaf axils and the bases of the leaves rather
the buds, young twigs, inflorescences, and
The hairiness
4
thunbergiana.
bisexual
true
and D.
species
sides.
specimens
patently hairy.
D.
I.,
Galapagos
rounded and
infraspeci-
the present
the materials from Asia,
Galapagos
polygamous (bisexual
is
& 28356 from Hawaii. This
may
donaea from
holds
seem
Madagascar,
from the
spatulata
var.
specimen,
In
nearly exclusively polygamous, mostly dioecious,
angustifolia,
specimen),
some
pistillode
same
anthers
(7 99,
andromonoecious. In the Pacific there is
the form called D. viscosa
the
the
in 9 flowers the
the
also
possibly
9,
Africa and
Tropical
are
area.
the Southafrican D.
d flowers
developed
area.
taxon, however.
be dioecious
to
probably
bisexual. In
also
may
direction, the flower conditions appear
comes
appeared
11
strongly reduced;
to
if these
even
fully
was
population
d and
for that
only
several
to
some
the gap between the present
change, widening
the American
to
geograph-
a
in the Hawaiian I., also
especially
infraspecific
any
eastern
an
show such
not
recognize
to
of these will be of value
from America in
turning
281
in distant parts of the
occur
may
be rather wide,
may
seems
from
recognizing
paper I have refrained
to
species of Dodonaea
differences in habitat. This
on
forms
comparable
fact that
range
even
least
at
in Australia. In these
fic
on
The variation in the leaves and in the fruits does
constant.
ical
Notes
to
densely,
sparsely hairy.
variable but
only rarely
conspicuous.
The variation of the leaves
ters, like
ten
length
much
and width,
narrower.
Some
linearis in South Africa
pa
from
Bourbon I.,
as
3—6.5
some
and
easily recognizable,
obovate leaves
(c.
7—10
x
2—3
x
are
etc.
cm,
the
one
cm,
c.
all kinds of characthe variation is of-
hand D.
ratio
population
characterized
ratio
regard
Locally,
c.
thunbergiana
5.5—21),
4)
with
a
from the
by large
var.
D. microcar-
with D. arizonica
Australia, comparable
the
hand,
being
on
0.3—0.75
forms from
from North America. On the other
constant
forms
extreme
(leaves
and
whole is wide and
may
a
ratio, texture, nervation,
Society
and
conspicuous
I. is rather
relatively
and
dense
broad
nerva-
tion.
In
the
regards
Hawaiian 1.
more
especially
the
variation is
characters. The
of the
fruit,
the
principal
probably
wider
characters
number of locules in
are
pistil
than
leaf
in
shape
any
and
other
size,
and fruit, size and
area
and
hairiness,
shape
of the
282
BLUMEA
and the width of the fruit
fruits,
viz. D. eriocarpa,
distinguished,
subdivided into
varieties and forms
about 65 specimens,
on
forms,
forms.
a
number of
formed
together
herent group in the
not
take
hardly
or
forms, appeared
identical with
paratype (Rock
one
type
stenoptera, takes
at
inflated with 3
comes
sible
group,
least
This
so
slightly hairy.
define
already
distinguishable
own;
to
as
form
that used
&
give
a
294.
Yunnan:
cit. 294.
IT.
INDIAN
(1972)
342,
463.
fig.
10846.
Lei
43932,
-
I.:
typical
clearly
or
but
clearly
co-
did
its many varieties and
D. sandwi-
variability.
identical collections
some
collected
mm
the
at
nearly
be
to
or
the wings broader,
inflated,
the top, and the
at
as
a
be-
body
hardly
pos-
recognize the varieties
to
even
whole
between D.
does
even
are
hardly
not
or
or
take
not
angustifolia
and D.
at
a
all
important literature and/or
some
L, M, RSA, SING, and U. If
under D.
viscosa
of the distribution of both
account
from FI,
are
are
of New Guinea.
separation
were
D.
connected with the
gradually
from the central group
most
is
locality,
same
wide; the body is glabrous but
incision
entities
Ming
in
396.
—
Taiwan:
Troup,
see
viscosa. The
Keng
Pradesh:
-
geographical
species,
selected
not
stated
sequence is
Panigrahi
Iran:
& Tawada
1
Liao
Sherff,
Arabia:
6549.
-
see
& Kuo
Mus.
Bonin I.:
Sherff,
2147,
Field
Nat. Hist.
Schimper 261,
(1978) 282, pi.
Kwantung:
1362,
see
-
66
Fujita&Shimizu
fig.
op. cit.
Bot.
266.
7
&
294.
Odashima
8.
-
—
Fukien:
Hainan:
65
see
Chun
17825, 17880; see
189.
Indian
Indian
-
1020/1.
Yunnan
Pakistan:
Brandis,
Silvic.
Fl.
188.
(1963) 494, fig.
India:
Madhya
Wu,
Amoy: Chung
-
SUBCONTINENT.
202;
946.329-153.
Griffith herb.
Fosberg 37873,Walker
see
Taiwan
-
L
in
Afghanistan:
-
Ryukyu
Woody
1
appeared
rather detailed
of the
num-
the sheets
glabrescens. Only the third 'species',
than
populations
Because of the
Groenhart
Iraq:
(1947)
JAPAN.
Tso
Li,
good
a
on
the Kew herbarium.
by
op.
the
specimens
some
to
CHINA.
Sherff,
and
the fruits become less
infraspecific
the literature citations
ORIENT.
Ser. 23
given
Even with the restricted material available it seemed
collections. The collections cited
otherwise
var.
no more
form
extreme
the citation
by
it
and of
position because of its peculiar fruits. These
extreme
an
from the inland
desirable
justifying
eriocarpa
Gradually,
clearcut
Distribution.
seems
atten-
varied collection of
variable,
with
and
214568)
few
species locally recognized
eriocarpa,
described. And the Hawaiian material
position of its
it
of D.
however.
shouldered
to
forms
in L
special
pay
was
a
As the
223—224).
rather uniform
the whole wide range of
s.n.
to
a
species
most
the
even
sandwicensis
4 rib-like wings
or
strongly glandular.
finally
of D.
the paratype
Heller 2871, the
central
D.
position.
nearly
were
delimited against each other
sharply
largest,
that
each
fell under the central group. Another member of this group,
available,
were
encompass
disposal
my
morphologically
were not
out
at
of all three
series with the
It turned
special
a
up
to
censis, of which
gradual
a
center.
1973:
of the identifications
Irrespective
species
encompassed
two
it seemed desirable
For this I had
coherent entities. Most of these entities
but
these characters three
the other
including representatives
sorted into
was
1983
and D. stenoptera. D. eriocarpa
sandwicensis,
varieties and
population.
ber of the varieties and
the material
wings. Mainly
2,
St. John, List Flow. PI. Hawaiian I.,
(see
the Hawaiian
to
No.
different from elsewhere
pattern appeared
tion
35
some
28,
D.
VOL.
-
Siddiqi
Trees
Trees
6616.
&
(1921)
-
Nasir
17;
see
Stewart,
(1906) 186; Talbot,
225.
Tamil Nadu:
-
6
Punjab:
colls.,
For.
Koelz
incl.
Cat.
Fl.
Fl. W.
Bombay
Pakistan
1
(1909)
4046, Schlagintweit
Hohenacker
1067,1068,
P.W. Lecnhouts: Notes
27984.
Shetty
Ceylon:
-
incl.
colls.,
7
extra-Australian
on
283
species ofDodonaea
Fosberg 50675, 50676;
Fl.
Trimen,
see
1
Ceyl.
(1893)
312.
INDO-CH1NA.
See
Fl.
Lecomte,
1
Indo-Chine
1005.
(1912)
Vietnam:
—
Chevalier
30454,
Poilane 976.
MALES1A.
Bali, Timor,
31
Java:
the
all
Vanoverbergh 1032,
&
AUSTRALIA.
Kanis
South
49
NSW
377;
see
Curtis
&
2
(1961) 29, pi.
&
43°30'
PACIFIC.
colls.,
on
incl.
New
246.
(1972)
Hotta
4750
partly.
Mus.
Bull.
Parks
6
Tahiti
and
(1961)
Flowers
and
109
(partly
as
2839.
A,
D.
herb.
Yoshida
995
island.
Maiden
-
35.
& Boor-
Tasmania:
-
Lord
-
New South
-
Bernice
-
P.
Bernice
NSW
Gunn
I.:
Howe
van
117587.
Tahiti
&
Mus.
6
colls.,
67
incl.
colls.,
eriocarpa); Degener,
128
incl.
Plants
in low
(1959)
Moore
Guam:
Rock, Indig.
see
189,
P.
Bishop
Setchcll &
Stone, Micro-
see
Herbst
Trees Hawaii
Fl. Hawaii
(1956, 1959)
PI.
(1961) pi.
as
D.
ed.
Tonga:
Degener 31549, 31550, 31551,
etc.;
the
176, at least
Bernice
8,
rev.
133. -
16
scrub
from
Fiji I.,
(1935)
220
—
Caledonia:
4253, 6593,
Brown,
see
10238.
Rarotonga: Philipson
—
Hawaii:
Smith
Bull.
Polynesia:
I.:
Society
Banks Penin-
common
Parham,
see
Mus. Bull.
Bishop
S.E.
—
L
(1967) 52, fig.
as
New
—
and most
A.C.
Group;
in
PI. Stud. ed. 2
2
ed.
far South
as
J.L.N,
9035,
131760.
L
in
Bishop
P.
186.
Colour,
in
reaches
Morrison
Hooker
Zeal. Native
New
Plants
Robinson
and the Malolo
(1970)
Raiatea.
—
55
in
Davies,
Degener 14299, 15347,
Yuncker,
see
Gomez
429:
Kanai, Oahu, Molokai, Lanai, Maui, Hawaii,
278, 281, pi.
1295.
13559,
Adams
(1975) 122, fig.
(1971) 549-554;
RSNH
Fl. Niue
62.
&
2619, 2837,
from
20391,
L.S. Smith 3549.
190,
2481, Haegi
Craven
1
2,
colls,
PNH
Territory: Chippendale
Dietrich
10034, 12577, 12744; mainly
incl.
163-164.
(1970) 396, fig.
from
824,
Chew
Levu,
1.;
ed.
Hoogland 6652,
Viti
Sykes,
see
9
Christophersen,
see
(1935)
from
I.:
1
Zealand
Bot.
colls.,
Levu,
Vava'u
1.:
130
547,
nesica
from
11
I.:
Vanua
Samoa:
-
Niue
—
Fiji
-
Group,
J.
Bernardi
3304,
Northern
-
2567,
Kraehenbuehl
Fosberg 30288,
New
Zealand
Hebrides:
18
17184,
1215.
New Zealand
Norfolk
-
Balansa
serpentine.
Yasawa
SL.
I.:
Sunda
Clemens
87 colls, from the whole
220574.
Blake
Streimann
Tasmania,
Zealand:
Fl.
3; Salmon,
New
123-125; Rivers,
c.
New
L
41747, Hartley,
incl.
Lesser
416,
Biinnemeijer 12093, Meijer 11049,
incl.
12605,
colls.,
Fl.
in
colls.,
-
F.ast.
Britton
New Guinea:
-
Mills
20
6
& Rudd
Allan,
see
Celebes:
-
peninsula.
NSW
Stud.
Philippines:
-
Eichler
Victoria:
Morris,
ZEALAND.
953.111-945;
incl.
Constable
—
Johnson
Balgooy 1136,
NEW
incl.
119215.
1.
Australia:
colls.,
and
West, Central,
Luzon.
Queensland:
-
21
colls.,
man
sula,
from
Western
1746.
Australia:
Wales:
from
Tanimbar
all from the southwestern
1376,
4950,
colls,
Babar and
(1913)
eriocarpa
and
D. sandwicensis.
TROPICAL
Lebrun
9
Beige
(1960)
(1928)
fig.
Schimper
&
ling
2
314,
705;
282.
892.
Zambia:
-
Angola:
-
Sudan
-
see
Sherff,
see
op.
2
cit.
Mozambique:
-
White,
Exell &
Jard. Bot.
282.
see
For. Fl.
19.
-
10242;
Mauritius:
SOUTH
Palmer &
&
see
Sieber 287.
AFRICA.
Pitman,
See
Trees
S.
Mem.
-
NORTH
6
Afr.,
AMERICA. Florida:
(1978)
Lippold,
op. cit.
op.
90,
as
89,
cit.
D.
as
84,
D.
as
bialata.
bialata.
D.
Mus.
Marloth,
2nd
arizonica.
S.
2
-
19.
2
2,
Steenis
2
-
126.
Congo
Sudan
Ethiopia:
-
Uganda:
see
Geesteranus
Exell,
Egge5033,
Fl. Zambesiaca
(1973)
see
d'Alleizette in
Bot.
158, pi.
1368-1370.
Fl.
and Central
44.
-
Malawi:
Sherff,
op.
cit.
91.
& Meeuse
(1925)
51
Rhodesia:
—
Madagascar:
B,
—
Maas
see
Congo Republic:
Hauman,
N.
494.
Africa:
224.
-
339, fig.
Moijamb.
(1954)
Hist. Nat. Ser.
(1972)
PI.
Kenya:
-
Fl.
(1962)
colls, incl. Lam
Afr.
(1952)
Tropical
Angol.
Flow.
Suppl. (1958)
Sousa,
ISLANDS.
Lippold,
New
Fl.
Natl.
5
ed.,
van
see
-
Fl.
Sudan
South
5.
(1948) 523, pi. 51;
Crowfoot,
Etat
N. Rhodesia
Reunion:
Goddijn 1685, Liitjeharms 7176,
tax.
—
Exell &
Mendon$a, Consp.
Capuron,
I
(1952) 378, pi.
MADAGASCAR and the MASCARENE
Decary
see
Anglo-Egypt.
Bull.
ed.
Uganda,
Ghana
Fl. Pare Albert
Republic:
PI.
Fl.
Cufodontis,
see
117.
(1966) 542, pi.
Stolz
see
Trees
Indig.
Irvine, Woody
see
Robijns,
see
39.
382, pi.
Tanganyika:
-
6088;
104(2); Andrews,
Dale,
5835.
Ghana:
AFRICA.
Malaisse
8141,
19
L951.64-405,
(1969) 27, pi.
4
fig.
12-
5284.
53:
Cape
B
(as
Prov.:
D.
thunbergiana);
9 colls, incl.
Lotsy
23916.
Wiss.
Mexico:
Beitr.
Friedr.
Bourgeau
California:
M.E.
312.
Jones
Schiller
-
Univ.
Arizona:
27087.
-
Jena Beitr.
Phyto-
Goodding 15567; see
Bermuda:
see
Lippold,
284
BLUMEA
CENTRAL
AMERICA.
and
Veracruz.
289.
D.
bialata.
27745.
27744,
heim
INDIES.
Lippold,
see
2970.
rina,
Smith
Grande
12140.
Peru
3
A
I.:
(1971) 751,
Uruguay:
at least
18
(1805)
Pflanzenr. 98
Fig.
3
H.
greenish,
glabrous
ous.
usually
Sepals
the
or
apex
more
to
not
lobed
to
in D.
cm,
in
2.
—
D.
Smith
&
Sao
Janeiro,
bialata.
189,
D.
23306,
Tiirck-
von
Breteler
see
de
Bruijn
1020.
Wiggins
Argentina:
-
Herter, Fl.
Illust.
&
see
Ann. Mis-
Schultes&
Ecuador:
-
Macbride,
see
Stcinbach
10707,
Krukoff
viscosa
Jacq.
in herb.
f.
44006,
Sta. Cata-
Croat,
20475, Cuatrecasas,
4097,
see
Rambo
32455,
Paulo, Parana,
Panama:
-
Cuatrecasas
Rudolphi
ratio
stiff
Fl.
Porter,
op. cit.
Sherff,
281,
-
I.
291.
2074.
Doming.
Jena Beitr.
elaeagnoides
Fl.
173.
Galapagos
Urug. (1942) fig.
Phy-
Radlk., Engl.
Willdenow 7515
at
7;
(B,
seen on
and
accordingly
beginning, terete,
Leaves obovate, 2.5-8
0.25 of the
length
sometimes either
brownish
varnished.
not
to
Inflorescences
the anther 1.5—1.8
mm
base
for
viscosa), mostly
in 9 flowers
partly 3-merous; style
body
4-7
mm
1.5—2.5
mm
monoeci-
completely
long,
mm
knob-
wide,
at
thin-pergamentaceous (slightly
mm,
conspicuously
not
4.5-5
the wings 2—4
high,
or
sometimes
mostly dioecious, rarely
long;
below
apiculate
Flowers unisexual,
Fruits:
style
most
at
viscosa:
from the
scurfy glandular.
nearly truncate,
sometimes
apex.
the
or
with D.
nearly
pergamentaceous, drying
glandular
or
comparison
sometimes
1.75-4.5, widest
sometimes
to
very
soon,
near
inland, up
the
glandular, glabrous, possibly
(acc.
to
Lippold, I.e.,
Bahamas, Cuba,
Eustatius
The
and the Florida
Sto.
(Arnoldo 3182, Boldingh
Desirade,
Grenada.
350
to c.
alt.;
m
acc.
to
Lippold (I.e.)
restricted
coast.
Florida
La
Fuertes
Ledeb. & Alderstam, Diss. Bot. PI.
facilitate the
NORTH AMERICA.
L),
as
263, Rutten-Pekelharing
Jamaica: Proctor
Wiss. Beitr. Friedr. Schiller Univ.
WEST INDIES.
St.
ex
see
fairly densely
or
coastal and
coral limestone
Distribution,
I.,
278,
90,
reddish.
Habitat:
to
as
de
Fiebrig 2501,
incl.
Type:
may
Pistil 2-merous,
stiff than
brownish
91,
galapagensis.
var.
map
variably glandular.
adnate
op. cit.
bialata.
Rio
Rojas 998,
colls.,
Lippold,
often
4. Stamens 6
slightly
op. cit.
Sherff,
Lippold,
Dombey 887, Fosberg 27696, 27703;
Rudolph
—
Curtiss
linearifolia.
Irwin, Harley
Bahia,
cit.
Cuba:
-
D.
D.
as
Colombia:
-
de
high .Branches
m
chartaceous
but
suppressed.
bed
9
viscosa
1371.
the apex; apex rounded
emarginate;
see
see
1 b.
1—2.25(-4)
x
op.
incl.
305.
resp.
90,
incl.
5040, 5229, 85049, 96921;
gnarly, greyish,
(—10)
D.
to
seen);
to
Nicaragua:
—
&
Tamanlipas, Hidalgo, Morelos,
British Honduras:
1040, 1539, 1914, 2334;
Werff
following diagnosis
Shrub up
rather
Bolivia:
(1933)
microfiche).
The
Benitez
colls.,
(1978) 95, pi. 1—5,
6
totax.
—
Lauber
181, 290, Stanford,
Domingo: Eggers 1885,
cit.
from
5.
cit.
Sto.
-
colls.,
11
Lippold,
see
elaeagnoides
(not
143,
Nuovo Leon,
bialata.
bialata,
Lippold, op.
see
6
-
as
D.
as
op.
D.
1333.
Brazil:
Sul;
der
Herter
as
(1977) 446, fig.
390.
van
Dodonaea
101,
57, 3971, 15022,
Peru:
-
Palmer
cit. 278.
op.
90,
Sherff,
see
88,
Venezuela:
—
(1956)
Galapagos
2.
do
23195.
Fosberg
-
& Klein
Bot. Gard. 63
op. cit.
Picarda
AMERICA.
Reitz
Rio
souri
cit.
Haiti:
-
Sherff,
see
Bahamas:
op.
incl.
colls.,
1983
2,
Durkee 75-63.
Puerto Rico:
-
SOUTH
44066,
Rica:
17
28, No.
Sonora, Chihuahua,
Lippold,
see
Costa
-
WEST
179;
Guatemala:
—
Honduras:
-
Mexico:
from
Taylor 2022, 2051, 2453,
VOL.
-
if
not
otherwise
mentioned).
Keys.
Domingo,
several
Puerto Rico
colls, in
U,
Stoffers
(P.
Sintenis
859),
the
Virgin
3503, Suringar several
colls,
P. W.
1. Dodonaea
Fig.
b.
D.
Ruth
Note.
van
a.
D.
a
s.n.
4
28, pi.
style (Sieber
in
s.n.
herb.
L
Fl.
285
Martinique 101).
905.136-157).
Both
x
-
2.
1—6.
fig.
—
different fruits
clearly
base
1
(fig.
b)
Abh.
are
—
allied with viscosa but
Radlk.,
1384;
style
me
sufficiently
Naturwiss. Vereine
Capuron,
for
Mem. Mus. Natl.
Bremen
smaller
strong
ar-
different.
8
(1884)
Hist. Nat. Ser.
J.M. Hillebrandt 3604,
Syntypes;
—
B,
Hilsenberg
(BM) (neither seen).
This species, endemic
not
the
to
good species,
madagascariensis
(1969)
species of Dodonaea
from the
style (Suringar
adnate
470; Engl. Pflanzenr. 98 (1933)
Bot. 19
free
wings
viscosa:
the unisexual flowers and the
wings apically
Dodonaea
extra-Australian
on
adnate to the
consider this
to
Notes
Crevel.
Especially
with the
guments
3.
fruits,
elaeagnoides: wings
Drawing by
and
Lecnhouts:
belong
to
the D.
to
Madagascar,
is the
only
viscosa group. It resembles
one
outside of Australia that does
mostly
D.
polyzyga
F. Muell. from
inland NW. Australia.
4.
Dodonaea
Brass 8379
(iso
in
This species
is
sepals (5
distinctly
4)
and
(Western Dist.,
Dodonaea viscosa
thor):
Merr. &
Perry,
J. Am. Arb.
Jacq.,
Sloane herb. v.97
allied
of
D.
to
stamens
viscosa;
D.
Enum.
(BM,
seen
else. Moreover,D. viscosa
elaeagnoides.
other
species
the
525.
(1940)
(12-15
In order
to
it
differs
instead of
Bensbach and Morehead
PI.
Syst.
on
meets
facilitate the
following diagnosis
is
(1760)
microfiche).
In America, the difference between D.
anywhere
21
—
Type:
L.J.
L).
instead of
New Guinea
5.
polyandra
—
19.
Fig.
an
comparison
exclusively
D.
the number of
to
SE.
and NE. Queensland.
Lectotype
—
1
in
It is restricted
5—9).
Subdists.)
viscosa and
here with
mainly
(present
au-
a.
angustifolia
other
closely
between D.
is slighter than
allied
species,
viz.
viscosa and the
two
based upon American material.
286
BLUMEA
Shrub
or
often
dular,
thicker,
terete
more
obovate,
5—12.5
above
sometimes in the
or
1.5-4.5
x
blackish,
cm,
2.25—3.75
so
in
either
just rounded,
nished
or
rarely
cous, scar
under the
1.66
Pistil
mm.
fruit
bisexual.
often
cleft
some
at
in
slightly
to
leaves
pergamenvarnished
not
leaves
some
flowers
some
not
var-
cadu-
5),
annular. Anthers 1.1-
nearly
to
several 3-merous
to
the apex. Fruits:
wide; membranous,
mm
coastal,
of distribution.
Only
mentioned. F.
and 9
some
B<j>rgesen
these
9; the
the
ones
body
on
9—13
the
mm
coloured, the body
straw
often 3-merous,
tifolia. Probably,
Distribution.
and for the
INDIAN
(1877)
the
it is
—
4121, from
Sumatra:
5
Quoi
colls.,
Coast, Benkulen,
neighbouring islands,
Kostermans
Ashton
-
wan,
S
the seeds
17829,
343,
Buwalda
and
clearly
female, the pisangus-
developed.
same
way
as
under D.
angustifolia
15
colls.,
6073,
incl.
the
Robinson
island,
New
-
1020/2;
75-1004, 76-659, both
incl.
see
Forest
Kurz,
from Chon
and
Java,
P.
from
SAN
Elmer
PI.
-
Malay
P.
Bawean.
Buri
Prov.,
80333,
Hou
31
Penang.
Java:
—
Lesser
551,
-
and
from
and Mindanao.
Southwest.
New Guinea:
New
11
from
Fl.
Burma
1
Ira I.
-
Viet-
-
25
I.:
Timor.
9
-
colls.,
from
colls.,
incl.
Borneo:
and
Sarawak, Sabah,
Merrill
Celebes:
Moluccas:
4
Sp.
colls.,
18
120,
Blanc.
incl.
colls.,
incl.
and
BW
Mapia
from
West
Thailand, Perak,
from
West Java
Iboet
8 colls,
601,
incl.
350,
incl.
Balambangan
from
Koorders
Morotai, Halmaheira,Ternate,
colls.,
Hanover,
-
colls,
Sunda
10452a, 12179, 16760,
and
2222, Lesger
Peninsula:
Sumba, Flores,
Rumph. 1,
Britain,
HB
Ajoeb 25, Diepenhorst
Krakatau.
North, Central,
and Ambon.
herb.
908.270-147.
L
Johore, Langkawi I.,
East
Dewol & Bindin
from
Buru, Ceram,
all over
is Curtiss
viscosa but for the
I.
Mindoro, Luzon, Polillo, Sibuyan,
Rachmat
also from Surinam.
specimen
the fruits resemble those of D.
well
are
Griffith
in
22071, Teijsmann 11177,
Philippines:
4755,
from
coll.
Maxwell
Phu
Dindings, Kelantan, Pahang,
and
clearly lobed,
as
Burma:
King's
Andamans:
MALESIA.
East
12702,
doubtful
only really
The distribution is given in the
INDO-CHINA. Thailand:
Coast,
is
6
some
same reason.
SUBCONTINENT.
287.
Pierre
style
hybrid
no
bisexual but has
partly bisexual, partly functionally
are
area
1091 from Surinam has the flowers
it resembles D.
Vegetatively
worldwide
the other should
or
Croix is monoecious with
mainly
is
(Sto. Domingo).
nearly
its
for Pons L.B.B.
true
alt.
m
character
one
Domingo
Hekking
viscosa. The
Cuba.
glandular leaves; the flowers
holds
250
at
throughout
different in
altitude.
m
D.
once
in U 24337B from St.
s.n.
same
undisputable
are
263 from Isla de Pinos,
nam:
uniform
specimens
250
at
occurs
partly functionally
til is
sandy beaches,
on
flowers. M. Fuertes 31 7 from Santo
6 flowers, and
All
mostly
present species is surprisingly
The
I.,
Leaves
widest
conspicuously glandular, glabrous.
Habitat;
I.
reticulately.
wide,
as
sometimes in
(not rarely
lobed
deeply
but
slightly
to
high, wings halfway (3.5—)4—6
not
be
long
glandular, accordingly
hardly
4
Sepals
from
varying
not
lengthwise
as
apiculate,
rounded and
mostly 2-merous,
specimen; style
same
times
Inflorescences
acute.
Flowers
so.
remaining reddish brown, sometimes
fissuring
nearly always inconspicuous,
usually
differ-
essentially
not
middle, thin-pergamentaceous (papyraceous
(only
or
1983
2,
uniform, nearly always hardly glan-
more
nearly always
drying greenish, glands
apex
but
and
taceous),
bud);
28, No.
Glabrous . Branchlets
high.
m
species,
American
somewhat
becoming
soon
1.5-3
rarely treelet,
from both other
ent
VOL.
-
Pala-
18818,
Bloembcrgen
Obi
and
Sula
2229, 3488, 12079, NGF 31703,
I.
P. W. Leenhouts:
AUSTRALIA.
Hubbard
PACIFIC.
lansa
157.
Northern
L.S.
3478,
1290
Marquesas: Gagne
viz.
Bikini
Atoll.
Deaw
184.
MADAGASCAR
chelles:
and
778
Jeffrey
6
WEST
INDIES.
Fuertes
Lippold,
I.e.
Lippold,
I.e.
see
Lippold,
this
see
St.
-
I.e.
I.e.
Curtiss
263;
I.e.
Eggers
Martin:
44
Grenada:
all
species
2243.
—
D. bialata,
Dodonaea arizonica
Jacq.
10
(1947)
(Sonora).
cm
peared
ring
x
extremely
in
2
I. coll.
from
incl. Ba-
colls.,
1965
from Guam.
293.
nr.
Marshall
—
-
I.: 4
and
Wotje, Likiep,
of D.
I.: Lam
&
Schiller
Univ.
I.e.'— Jamaica:
senegalensis.
Meeuse6577.
Sey-
—
Jena Beitr.
Phyto-
2904,
3031
Lippold,
in
Lippold,
see
3743.
L
I.e.
Trinidad:
-
I.e.
in U
Saint Croix:
-
B, Suringar
3627;
see
Santo Do-
-
4507, Sintenis 108, Wagner 442;
A.
295.
—
colls.
Brazil:
-
Martius
accepted
either
author,
are
24337
905.136-159.
—
Sieber
8996.
sec
see
Antigua:
-
Martinique:
Broadway
B;
101;
Tobago:
-
mm,
c.
Sudamer.
Friedr.
Beitr.
Nelson,
Amer.
Jena Beitr.
nearly
as
J.
conditions
phenotypically
in
5
Schiller
D.
to
Bot.
J. Bot. 32
(1933)
or
by
some
(1937)
55;
Fl. Illust.
Urug.
Univ.
Beitr.
Jena
6
Phytotax.
angustifolia.
21
6
(1934)
576; H.
(RM,
Wiss.
Lippold,
map 1.
(1978) 82, pi. 1-5,
(1945)
A. Nelson 11276
a
shrub
as
seems
long
to
as
no
-
D.
214; Field Mus. Natl. Hist.
not
be
an
than
more
wide).
seen).
collection,
occur
form of D.
173,
occurrence
genetically
high
with
taxa.
least
by
(up
me
ap-
occur-
under drier
con-
from Bolivia; other
comparable
strengthens
defined
seen
thunbergiana
of
at
narrow
angustifolia,
growing
is known
in South Africa (D.
different parts of the world
rather than
m
The few specimens
extreme
Steinbach
1.5
and the leaves very
other American Dodonaea and
southern Australia. The
extreme
Bahia.
from
occurring in the U.S.A. (Arizona, Texas) and Mexi-
by being
narrow-leaved forms
these
I.e.
Lippold,
Radlkofer
by
Bot.
Phytotax.
Amer.
Type;
any
sec
mentioned.
10 times
identical
397;
discussed.
fairly densely minutely patent hairy
Central and
are
Comoro
Rico: Heller
the present author
Sherff (1947)
farther North than
A
Niue
460
Wiss. Beitr. Friedr.
B,
Revista
Wiss.
by
be dioecious. This
to
ditions.
and
4—8
2504
names,
not
Characterized
the young parts
9
incl
REJECTED SPECIES NAMES
f. arizonica Sherff,
Mentioned by
to
New Caledonia:
—
Sykes
908.269-1219, Type
L
210, Broadway
Herter,
Type:
by Lippold,
to
Bot. Ser. 23
co
colls.,
from Praslin.
Puerto
the present
rejected by
Beitr. Friedr. Schiller Univ.
viscosa
in
from St. Thomas.
Duss
Beard
Surinam:
Dodonaea arborea
Reduced
2.
5
Queensland:
-
9830.
chapter
(1978) 86,
I.:
McGregor
Lippold,
see
-
Boldingh
Guadeloupe:
—
-
later author but
1.
Niue
-
ISLANDS.
Lippold,
Lippold,
1.:
AMERICA.
(1942) fig.
I.
306.
Procter 3948
IV.
In
Guadalcanal.
Marianas:
—
Warnecke
Florida:
see
Virgin
Webster & Miller
SOUTH
Wessel
21799, 22015, Taylor 46-1095, 46-1442,
Mahe,
Cuba:
317;
-
Lippold,
see
from
287
106.
(1978)
mingo:
from
Savaii.
the MASCARENE
from
NORTH AMERICA.
tax.
2458
from
I.
Eiao
Togo:
-
3282
Senegal: Anonymus
AFRICA.
Sp.
356
from
St.John & Cowan
TROPICAL
Ghana:
Latz
species ofDodonaea
12525.
Kajewski
Reinecke
extra-Australian
on
Territory:
11013,
I.:
Solomon
Samoa:
-
colls.,
Smith
Notes
var.
linearis)
forms under
the impression that
288
3.
BLUMF.A
Dodonaea bialata
Wiss. Beitr. Friedr.
—
H.,
Humboldt &
Type:
B. & K.,
Schiller Univ.
VOL.
1983
2,
Nov. Gen.
Sp.
Jena Beitr.
Phytotax.
355
Bonpland
No.
28,
herb.
=
5
H.
(1821) 104, pi. 442;
6
Lippold,
(1978) 85, pi. 1—5,
Willdenow 7316
(B, P;
seen
map 1.
micro-
on
fiche).
For
description
a
referred
be
may
the
Chapter III,
to
of American D.
diagnosis
angustifolia.
Reduced
4.
Radlkofer
by
author
present
Dodonaea
D.
to
eriocarpa
38
(1951) 57-61;
Flow. PI.
See
Reduced
5.
D.
to
Dodonaea
viscosa
Jacq.
1369.
f. burmanniana
the
Radlk., by
the discussion
Menzies
D.
on
20
(1949) 4-7;
Fl. Hawaii
Jacq.
viscosa
Amer. J. Bot. 32
(1809); Sherff,
Mus.
Bishop
D.
—
Type:
—
12
Cyclop.
P.
(1953) 15; Degener,
223.
(1975)
1.
(1933)
Chapter III,
Bernice
Pap.
Bot. Leafl. 8
Hawaiian
Pflanzenr. 98
D.
to
Smith in Rees,
Occas.
(1945) 204-213;
(1933)
angustifolia
Amer. J. Bot.
List
(1956); St.John,
subf. eriocarpa
Radlk.,
Engl.
(not seen).
angustifolia.
angustifolia
lagunensis
M.E. Jones, Contrib. West. Bot. 18 (1933)
57.
—
Type:
M.E.
Jones 27087 (RSA).
Apparently,
I
273),
who
seemed
coastal
plant
D.
under D.
Jones'
at
first
c.
4—7
only,
original
f.
author,
every
repanda
ecologically improbable.
as
but unknown
Tropics,
These arguments
that D.
even
Radlk. This reduction
the contrary, is described from the
well
as
lagunensis
of D.
description
worthwhile
and leaves
mainly
beneath
glands
the
the conclusion
seems
1.75—2
x
well
Subtropics.
angular, brown,
middle,
to
Jacq.
nearly
by Lip-
Mus. Nat. Hist. Bot. Ser. 23,
is
a
D.
to
viscosa
1947:
true
s.s.
D.
is
a
from the American West
Mts. in
Laguna
as
a
California,
of the type col-
study
angustifolia
synonym of D.
viscosa.
Twigs, petioles,
midrib
on
the
to
me
rather than of D.
characters it
as
by
Sherff (Field
by
viscosa
and is restricted
lagunensis,
lection led
has been overlooked
mentioned
geographically
i.e. inland and in
As
name
only
cited it
viscosa
coast.
this
found it
pold.
cm,
and
to
give
least
a
short
on
the
midrib,
very
about 2.25—3.5 times
especially
on
long
the
diagnosis
sparsely
soon
as
only
very
on
becoming
as
wide,
upper
side,
few essential
based upon the type:
the latter above
thinly shortly hairy,
slightly glandular,
conspicuous
includes
lagunensis
at
on
the base of the
the minor veins.
terete
widest
Twigs
and black. Leaves
slightly
leaves
above the
varnished;
apex
rounded, apiculate. Inflorescences short and few-flowered, hardly glandular. Flowers
unisexual
probably
and
the
body
7.5
6.
c.
mm
scar
1
of the
cm
wide,
Dodonaea
as
under the fruit the
calyx
is annular.
scars
of the
stamens
Sepals 4, partly persistent.
high, distinctly glandular
and very
are
not
conspicuous
Fruits 2-merous, the
sparsely short-hairy,
the
wings
up
to
sometimes reddish.
linearifolia
(1858)
407; H. Lippold,
(1978)
100, pi.
1-5,
Linden
Wiss.
map 2.
—
ex
Turcz., Bull. Soc.
Beitr. Friedr.
Type:
Schiller
Imp.
Univ.
Naturalistes Moscou 31
Jena Beitr.
Phytotax.
Linden 2070 (KW; iso in A, G, NY;
not
6
seen).
P. W.
Reduced
This
7.
Radlkofer
by
seems to
Lecnhouts:
be
Reduced
(NY;
by
(1933:
224.
—
See
D.
9.
Fl.
Degener,
Hawaii
Chapter
D.
viscosa
Jacq.
289
angustifolia
var.
Benth.
angustifolia
-
Small, Cuthbert &
Type:
(1978:
95)
D.
to
Ledeb. &
elaeagnoides
Alderstam. This
the present author.
by
(1959)
Jules Remy 567
Type:
to
Small, Torreya 25 (1925) 39.
8. Dodonaea sandwicensis Sherff,
6;
1366)
species of Dodonaea
seen).
not
Lippold
is shared
opinion
extra-Australian
on
of the narrow-leaved forms of D.
one
Dodonaea microcarya
Matthaus 9105
Notes
the
III,
Amer. J.
with
(P;
discussion
Bot. 32
fig.; St.John,
(1945) 202;
Bot. Leafl. 9
(1954)
List Flow. PI.
Hawaiian I.
(1973)
seen).
not
the
on
of D.
variability
Reduced
angustifolia.
to
angustifolia
Dodonaea stenoptera Hillebr., Fl. Hawaiian Isl.
98
(1933)
waiian I.
(1973)
Among D.
the
most
Amer. J.
1374; Sherff,
224.
Type:
-
angustifolia
Hillebrand (B;
the broad
(1888) 88; Radlk., Engl.
(1945) 213; St.John,
not
deviating
no
more
than
1
—
1.5
kind of fruit
extreme
mm
to
the
List
Pflanzenr.
Flow.
PI.
Ha-
seen).
accepted by
sense
form because of its fruits. These
with rib-like wings
sition from the
in
Bot. 32
are
the present
author, this is
inflated, slightly horny,
broad. However, there is
a
'normal' less-inflated and
gradual
and
tran-
broad-winged
fruits.
To be reduced
10.
to
D.
angustifolia
Dodonaea thunbergiana Ecklon &
Marloth,
569.
-
D.
viscosa
Jacq.
Ecklon <£
Type:
D.
Fl. S. Afr. 2, 2
Zeyher,
(1925) 158, pi. 53;
f. thunbergiana
Zeyher
419
(iso
in
Enum. PL Afric. Austral.
Adamson & Salter, Fl.
Radlk., Engl.
Cape
Pflanzenr. 98
(1933)
(1950)
1372.
—
1
(1860)
242.
—
Drège (not seen).
Especially
the
var.
and D.
linearifolia
6.5
x
cm
(1835) 54;
Pen.
L).
thunbergiana var. linearis Sonder in Harvey & Sonder, Fl. Cap.
Type:
1
3—7.5
mm, c.
To be reduced
linearis has
very
narrow
leaves, comparable
with D. arizonica
in America and with certain Australian forms. These leaves
to
D.
5.5—21
times as
angustifolia.
long
as
wide.
are
3—