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Essentials of Human Anatomy & Physiology Seventh Edition Elaine N. Marieb Chapter 8 Special Senses Slides 8.1 – 8.54 Lecture Slides in PowerPoint by Jerry L. Cook Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings The Senses General senses of touch Temperature Pressure Pain Special senses Smell Taste Sight Hearing Equilibrium Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings •Special sense receptors are either large, complex sensory organs (eyes and ears) or localized clusters or receptors (taste buds and olfactory epithelium) Slide 8.2 The Eye and Vision 70 percent of all sensory receptors are in the eyes Each eye has over a million nerve fibers Protection for the eye Most of the eye is enclosed in a bony orbit A cushion of fat surrounds most of the eye Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.3 Accessory Structures of the Eye Eyelids Eyelashes Meibomian glands – modified sebaceous glands produce an oily secretion to lubricate the eye Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Figure 8.1b Slide 8.4 Accessory Structures of the Eye Ciliary glands – modified sweat glands between the eyelashes Conjunctiva Membrane that lines the eyelids Connects to the surface of the eye Secretes mucus to lubricate the eye Figure 8.1b Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.5 Accessory Structures of the Eye Lacrimal apparatus Lacrimal gland – produces lacrimal fluid Lacrimal canals – drains lacrimal fluid from eyes Figure 8.1a Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.6 Accessory Structures of the Eye Lacrimal sac – provides passage of lacrimal fluid towards nasal cavity Nasolacrimal duct – empties lacrimal fluid into the nasal cavity Figure 8.1a Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.7 Function of the Lacrimal Apparatus Properties of lacrimal fluid Dilute salt solution (tears) Contains antibodies and lysozyme Protects, moistens, and lubricates the eye Empties into the nasal cavity Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.8 Extrinsic (External) Eye Muscles Muscles attach to the outer surface of the eye Produce gross eye movements and make it possible to follow moving objects Figure 8.2 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.9 Structure of the Eye The wall is composed of three tunics – coats while the interior is filled with fluids called humors Fibrous tunic (sclera)– outside layer Choroid – middle layer Sensory tunic – inside layer Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Figure 8.3a Slide 8.10 The Fibrous Tunic Sclera White connective tissue layer Seen anteriorly as the “white of the eye” Cornea Transparent, central anterior portion Allows for light to pass through Repairs itself easily The only human tissue that can be transplanted without fear of rejection – no blood vessels Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.11 Choroid Layer Blood-rich nutritive tunic Dark pigment prevents light from scattering inside the eye Modified interiorly into two structures Ciliary body – smooth muscle to which the lends is attached Iris Pigmented layer that gives eye color Pupil – rounded opening in the iris for light to enter Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.12 Sensory Tunic (Retina) Contains receptor cells (photoreceptors) Rods Cones Signals pass from photoreceptors via a two-neuron chain Bipolar neurons Ganglion cells Signals leave the retina toward the brain through the optic nerve Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.13 Neurons of the Retina Figure 8.4 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.14 Neurons of the Retina and Vision Rods Most are found towards the edges of the retina Allow dim light vision and peripheral vision Perception is all in gray tones Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.15 Neurons of the Retina and Vision Cones Allow for detailed color vision in bright light Densest in the center of the retina Fovea centralis – area of the retina with only cones – lateral to each blind spot No photoreceptor cells are at the optic disk, or blind spot – where the optic nerve leaves the eyeball Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.16 Cone Sensitivity There are three types of cones Different cones are sensitive to different wavelengths Color blindness is the result of lack of one cone type Figure 8.6 Impulses received at the same time are interpreted as intermediate colors Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.17 Lens Biconvex crystal-like structure Held in place by a suspensory ligament attached to the ciliary body Figure 8.3a Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.18 Internal Eye Chamber Fluids Aqueous humor Watery fluid found in chamber between the lens and cornea Similar to blood plasma Helps maintain intraocular pressure Provides nutrients for the lens and cornea Reabsorbed into venous blood through the canal of Schlemm Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.19 Internal Eye Chamber Fluids Vitreous humor Gel-like substance behind the lens Keeps the eye from collapsing inward by reinforcing it internally Lasts a lifetime and is not replaced Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.20 Internal Eye Chamber Fluids Cataracts form when the lens becomes increasingly hard and opaque over time requiring a transplant or special glasses Glaucoma results when the drainage of aqueous humor is blocked and pressure within the eye increases dramatically and compresses the delicate retina and optic nerve causing pain and blindness Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.21 Lens Accommodation Light must be focused to a point on the retina for optimal vision – done by the lens The resting eye is set for distance vision (over 20 ft away) The lens must change shape to focus for closer objects – accommodation Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Figure 8.9 Slide 8.22 Images Formed on the Retina Figure 8.10 •Real image (reversed left to right, and upside down) formed on the retina Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.23 Visual Pathway Photoreceptors of the retina Optic nerve Optic nerve crosses at the optic chiasma to the opposite side Fiber tracts that result are the optic tracts Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Figure 8.11 Slide 8.24 Visual Pathway Optic tracts contains fibers from the lateral side of the eye on the same side and the medial side of the opposite eye The optic tract fibers synapse with neurons in the thalamus, whose axons form optic radiation Visual cortex of the occipital lobe Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Figure 8.11 Slide 8.25 Eye Reflexes Internal muscles are controlled by the autonomic nervous system Bright light causes pupils to constrict through action of radial and ciliary muscles – photopupillary reflex Viewing close objects causes accommodation – accommodation pupillary reflex External muscles control eye movement to follow objects Viewing close objects causes convergence (eyes moving medially (toward nose)) Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.26 The Ear Houses two senses Hearing Equilibrium (balance) Receptors are mechanoreceptors – respond to physical forces Although these two sense organs are housed together in the ear, their receptors respond to different stimuli and are activated independently Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.27 Anatomy of the Ear The ear is divided into three areas Outer (external) ear Middle ear Inner (internal) ear Figure 8.12 Outer and middle ear structures involved in hearing only while inner ear functions in both equilibrium and hearing Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.28 The External Ear Structures of the external ear Pinna (auricle) – the ear External auditory canal Figure 8.12 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.29 The External Auditory Canal Narrow chamber in the temporal bone Lined with skin Ceruminous (wax) glands are present that secrete earwax – cerumen Ends at the tympanic membrane (eardrum) where sound waves hit and cause vibrations Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.30 The Middle Ear or Tympanic Cavity Air-filled cavity within the temporal bone Two tubes are associated with the inner ear – the oval window and the inferior, membranecovered round window The opening from the auditory canal is covered by the tympanic membrane The auditory tube connecting the middle ear with the throat Allows for equalizing pressure during yawning or swallowing – This tube is otherwise collapsed Slide 8.31 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Bones of the Tympanic Cavity Three bones span the cavity – the ossicles Malleus (hammer) Incus (anvil) Stapes (stirrup) Figure 8.12 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.32 Bones of the Tympanic Cavity Vibrations from eardrum move the malleus These bones transfer sound to the inner ear Figure 8.12 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.33 Inner Ear or Bony Labrynth A maze of bony chambers within the temporal bone called the osseous or bony labyrinth Cochlea Vestibule Semicircular canals Filled with a plasma-like fluid called perilymph Inside is a membranous labyrinth that contains a thicker fluid called endolymph Figure 8.12 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.34 Organs of Hearing Organ of Corti Located within the cochlea Receptors = hair cells on the basilar membrane – hearing receptors Gel-like tectorial membrane is capable of bending hair cells Cochlear nerve attached to hair cells transmits nerve impulses to auditory cortex on temporal lobe Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.35 Organs of Hearing Figure 8.13 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.36 Mechanisms of Hearing Vibrations from sound waves move tectorial membrane Hair cells are bent by the membrane An action potential starts in the cochlear nerve Continued stimulation can lead to adaptation – stop responding to those sounds Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.37 Mechanisms of Hearing Figure 8.14 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.38 Organs of Equilibrium Receptor cells are in two structures Vestibule Semicircular canals Figure 8.16a, b Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.39 Organs of Equilibrium Equilibrium has two functional parts Static equilibrium Dynamic equilibrium Figure 8.16a, b Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.40 Static Equilibrium Maculae – receptors in the vestibule Report on the position of the head with respect to gravity – help determine up from down Send information via the vestibular nerve Anatomy of the maculae Hair cells are embedded in the otolithic membrane Otoliths (tiny stones) float in a gel around the hair cells Movements cause otoliths to bend the hair cells which send impulses along the vestibular nerve to the cerebellum Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.41 Function of Maculae Figure 8.15 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.42 Dynamic Equilibrium Crista ampullaris – receptors in the semicircular canals Tuft of hair cells Cupula (gelatinous cap) covers the hair cells Figure 8.16c Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.43 Dynamic Equilibrium Receptors respond to angular or rotatory movements of the head Action of angular head movements The cupula stimulates the hair cells – gelatinous cap An impulse is sent via the vestibular nerve to the cerebellum Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Figure 8.16c Slide 8.44 Chemical Senses – Taste and Smell Both senses use chemoreceptors Stimulated by chemicals in solution Taste has four types of receptors Smell can differentiate a large range of chemicals Both senses complement each other and respond to many of the same stimuli Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.45 Olfaction – The Sense of Smell Olfactory receptors are in the roof of the nasal cavity Neurons with long cilia – olfactory hairs Chemicals must be dissolved in mucus for detection Impulses are transmitted via the olfactory filaments which makes up the olfactory nerve Interpretation of smells is made in the olfactory cortex Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.46 Olfactory Epithelium Figure 8.17 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.47 The Sense of Taste Taste buds house the receptor organs Location of taste buds Most are on the tongue Soft palate Inner cheeks Figure 8.18a, b Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.48 The Tongue and Taste The dorsal tongue is covered with projections called papillae Filiform papillae – sharp with no taste buds Fungiform papillae – rounded with taste buds Circumvallate papillae – large papillae with taste buds Taste buds are found on the sides of papillae Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.49 Structure of Taste Buds The specific cells that respond to chemical dissolved in saliva are epithelial cells Gustatory cells are the receptors Surrounded by supporting cells in the taste bud Have gustatory hairs (long microvilli) Hairs are stimulated by chemicals dissolved in saliva Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.50 Structure of Taste Buds Impulses are carried to the gustatory complex by several cranial nerves because taste buds are found in different areas Facial nerve – anterior tongue specifically Glossopharyngeal nerve Vagus nerve Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.51 Anatomy of Taste Buds Figure 8.18 Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.52 Taste Sensations Sweet receptors Sugars •May respond to the OH Saccharine Some amino acids Sour receptors Acids May respond to the H+ Bitter receptors Alkaloids Salty receptors Metal ions in solution Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.53 Developmental Aspects of the Special Senses Formed early in embryonic development Eyes are outgrowths of the brain All special senses are functional at birth Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings Slide 8.54