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Essentials of Human Anatomy & Physiology
Seventh Edition
Elaine N. Marieb
Chapter 8
Special Senses
Slides 8.1 – 8.54
Lecture Slides in PowerPoint by Jerry L. Cook
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
The Senses
 General senses of touch
 Temperature
 Pressure
 Pain
 Special senses
 Smell
 Taste
 Sight
 Hearing
 Equilibrium
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
•Special sense
receptors are either
large, complex
sensory organs
(eyes and ears) or
localized clusters or
receptors (taste
buds and olfactory
epithelium)
Slide 8.2
The Eye and Vision
 70 percent of all sensory receptors are
in the eyes
 Each eye has over a million nerve fibers
 Protection for the eye
 Most of the eye is enclosed in a bony orbit
 A cushion of fat surrounds most of the eye
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.3
Accessory Structures of the Eye
 Eyelids
 Eyelashes
 Meibomian glands
– modified
sebaceous
glands
produce an
oily secretion
to lubricate
the eye
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 8.1b
Slide 8.4
Accessory Structures of the Eye
 Ciliary glands – modified sweat glands
between the eyelashes
 Conjunctiva
 Membrane that
lines the eyelids
 Connects to the
surface of the eye
 Secretes mucus
to lubricate the eye
Figure 8.1b
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.5
Accessory Structures of the Eye
 Lacrimal
apparatus
 Lacrimal gland –
produces lacrimal
fluid
 Lacrimal canals –
drains lacrimal
fluid from eyes
Figure 8.1a
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.6
Accessory Structures of the Eye
 Lacrimal sac –
provides passage
of lacrimal fluid
towards nasal
cavity
 Nasolacrimal duct
– empties lacrimal
fluid into the nasal
cavity
Figure 8.1a
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.7
Function of the Lacrimal Apparatus
 Properties of lacrimal fluid
 Dilute salt solution (tears)
 Contains antibodies and lysozyme
 Protects, moistens, and lubricates the
eye
 Empties into the nasal cavity
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Slide 8.8
Extrinsic (External) Eye Muscles
 Muscles attach to the outer surface of the
eye
 Produce gross eye movements and make it
possible to follow moving objects
Figure 8.2
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.9
Structure of the Eye
 The wall is composed of three tunics – coats
while the interior is filled with fluids called humors
 Fibrous tunic (sclera)–
outside layer
 Choroid –
middle
layer
 Sensory
tunic –
inside
layer
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 8.3a
Slide 8.10
The Fibrous Tunic
 Sclera
 White connective tissue layer
 Seen anteriorly as the “white of the eye”
 Cornea
 Transparent, central anterior portion
 Allows for light to pass through
 Repairs itself easily
 The only human tissue that can be transplanted
without fear of rejection – no blood vessels
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.11
Choroid Layer
 Blood-rich nutritive tunic
 Dark pigment prevents light from scattering
inside the eye
 Modified interiorly into two structures
 Ciliary body – smooth muscle to which the lends
is attached
 Iris
 Pigmented layer that gives eye color
 Pupil – rounded opening in the iris for light to
enter
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.12
Sensory Tunic (Retina)
 Contains receptor cells (photoreceptors)
 Rods
 Cones
 Signals pass from photoreceptors via a
two-neuron chain
 Bipolar neurons
 Ganglion cells
 Signals leave the retina toward the brain
through the optic nerve
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Slide 8.13
Neurons of the Retina
Figure 8.4
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Slide 8.14
Neurons of the Retina and Vision
 Rods
 Most are found towards the edges of the
retina
 Allow dim light vision and peripheral vision
 Perception is all in gray tones
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.15
Neurons of the Retina and Vision
 Cones
 Allow for detailed color vision in bright light
 Densest in the center of the retina
 Fovea centralis – area of the retina with
only cones – lateral to each blind spot
 No photoreceptor cells are at the optic
disk, or blind spot – where the optic
nerve leaves the eyeball
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.16
Cone Sensitivity
 There are three types of
cones
 Different cones are
sensitive to different
wavelengths
 Color blindness is the result
of lack of one cone type
Figure 8.6
 Impulses received at the
same time are interpreted
as intermediate colors
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Slide 8.17
Lens
 Biconvex crystal-like structure
 Held in place by a suspensory ligament
attached to the ciliary body
Figure 8.3a
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Slide 8.18
Internal Eye Chamber Fluids
 Aqueous humor
 Watery fluid found in chamber between the
lens and cornea
 Similar to blood plasma
 Helps maintain intraocular pressure
 Provides nutrients for the lens and cornea
 Reabsorbed into venous blood through the
canal of Schlemm
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Slide 8.19
Internal Eye Chamber Fluids
 Vitreous humor
 Gel-like substance behind the lens
 Keeps the eye from collapsing inward by
reinforcing it internally
 Lasts a lifetime and is not replaced
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.20
Internal Eye Chamber Fluids
 Cataracts form when the
lens becomes increasingly
hard and opaque over time
requiring a transplant or
special glasses
 Glaucoma results when the drainage of
aqueous humor is blocked and pressure
within the eye increases dramatically and
compresses the delicate retina and optic
nerve causing pain and blindness
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.21
Lens Accommodation
 Light must be focused
to a point on the retina
for optimal vision –
done by the lens
 The resting eye is set
for distance vision
(over 20 ft away)
 The lens must change
shape to focus for
closer objects –
accommodation
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Figure 8.9
Slide 8.22
Images Formed on the Retina
Figure 8.10
•Real image (reversed left to right, and
upside down) formed on the retina
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.23
Visual Pathway
 Photoreceptors of
the retina
 Optic nerve
 Optic nerve crosses
at the optic chiasma
to the opposite side
 Fiber tracts that
result are the optic
tracts
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 8.11
Slide 8.24
Visual Pathway
 Optic tracts contains fibers
from the lateral side of the
eye on the same side and
the medial side of the
opposite eye
 The optic tract fibers
synapse with neurons in
the thalamus, whose axons
form optic radiation
 Visual cortex of the
occipital lobe
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Figure 8.11
Slide 8.25
Eye Reflexes
 Internal muscles are controlled by the
autonomic nervous system
 Bright light causes pupils to constrict through
action of radial and ciliary muscles –
photopupillary reflex
 Viewing close objects causes accommodation
– accommodation pupillary reflex
 External muscles control eye movement to
follow objects
 Viewing close objects causes convergence
(eyes moving medially (toward nose))
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.26
The Ear
 Houses two senses
 Hearing
 Equilibrium (balance)
 Receptors are mechanoreceptors –
respond to physical forces
 Although these two sense organs are
housed together in the ear, their receptors
respond to different stimuli and are
activated independently
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.27
Anatomy of the Ear
 The ear is divided into
three areas
 Outer (external)
ear
 Middle
ear
 Inner (internal)
ear
Figure 8.12
 Outer and middle ear structures involved in
hearing only while inner ear functions in both
equilibrium and hearing
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.28
The External Ear
 Structures of
the external ear
 Pinna (auricle) –
the ear
 External
auditory canal
Figure 8.12
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.29
The External Auditory Canal
 Narrow chamber in the temporal bone
 Lined with skin
 Ceruminous (wax) glands are present
that secrete earwax – cerumen
 Ends at the tympanic membrane
(eardrum) where sound waves hit and
cause vibrations
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.30
The Middle Ear or Tympanic Cavity
 Air-filled cavity within the temporal bone
 Two tubes are associated with the inner ear
– the oval window and the inferior, membranecovered round window
 The opening from the auditory canal is
covered by the tympanic membrane
 The auditory tube connecting the middle ear
with the throat
 Allows for equalizing pressure during yawning
or swallowing – This tube is otherwise
collapsed
Slide 8.31
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Bones of the Tympanic Cavity
 Three bones span
the cavity – the
ossicles
 Malleus (hammer)
 Incus (anvil)
 Stapes (stirrup)
Figure 8.12
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.32
Bones of the Tympanic Cavity
 Vibrations from
eardrum move
the malleus
 These bones
transfer sound
to the inner ear
Figure 8.12
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Slide 8.33
Inner Ear or Bony Labrynth
 A maze of bony chambers within the temporal
bone called the osseous or bony labyrinth
 Cochlea
 Vestibule
 Semicircular
canals
 Filled with a plasma-like fluid called perilymph
 Inside is a membranous labyrinth that contains
a thicker fluid called endolymph
Figure 8.12
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Slide 8.34
Organs of Hearing
 Organ of Corti
 Located within the cochlea
 Receptors = hair cells on the basilar
membrane – hearing receptors
 Gel-like tectorial membrane is capable of
bending hair cells
 Cochlear nerve attached to hair cells
transmits nerve impulses to auditory cortex
on temporal lobe
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Slide 8.35
Organs of Hearing
Figure 8.13
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Slide 8.36
Mechanisms of Hearing
 Vibrations from sound waves move
tectorial membrane
 Hair cells are bent by the membrane
 An action potential starts in the cochlear
nerve
 Continued stimulation can lead to
adaptation – stop responding to those
sounds
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Slide 8.37
Mechanisms of Hearing
Figure 8.14
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Slide 8.38
Organs of Equilibrium
 Receptor cells are in two structures
 Vestibule
 Semicircular canals
Figure 8.16a, b
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Slide 8.39
Organs of Equilibrium
 Equilibrium has two functional parts
 Static equilibrium
 Dynamic equilibrium
Figure 8.16a, b
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Slide 8.40
Static Equilibrium
 Maculae – receptors in the vestibule
 Report on the position of the head with respect to
gravity – help determine up from down
 Send information via the vestibular nerve
 Anatomy of the maculae
 Hair cells are embedded in the otolithic membrane
 Otoliths (tiny stones) float in a gel around the hair
cells
 Movements cause otoliths to bend the hair cells
which send impulses along the vestibular nerve to
the cerebellum
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Slide 8.41
Function of Maculae
Figure 8.15
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Slide 8.42
Dynamic Equilibrium
 Crista ampullaris –
receptors in the
semicircular canals
 Tuft of hair cells
 Cupula (gelatinous cap)
covers the hair cells
Figure 8.16c
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Slide 8.43
Dynamic Equilibrium
 Receptors respond to angular or rotatory
movements of the head
 Action of angular head
movements
 The cupula stimulates the
hair cells – gelatinous cap
 An impulse is sent via the
vestibular nerve to the cerebellum
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Figure 8.16c
Slide 8.44
Chemical Senses – Taste and Smell
 Both senses use chemoreceptors
 Stimulated by chemicals in solution
 Taste has four types of receptors
 Smell can differentiate a large range of
chemicals
 Both senses complement each other
and respond to many of the same
stimuli
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.45
Olfaction – The Sense of Smell
 Olfactory receptors are in the roof of the
nasal cavity
 Neurons with long cilia – olfactory hairs
 Chemicals must be dissolved in mucus for
detection
 Impulses are transmitted via the olfactory
filaments which makes up the olfactory nerve
 Interpretation of smells is made in the
olfactory cortex
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Slide 8.46
Olfactory Epithelium
Figure 8.17
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Slide 8.47
The Sense of Taste
 Taste buds
house the
receptor
organs
 Location of
taste buds
 Most are on
the tongue
 Soft palate
 Inner cheeks
Figure 8.18a, b
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Slide 8.48
The Tongue and Taste
 The dorsal tongue is covered with
projections called papillae
 Filiform papillae – sharp with no taste buds
 Fungiform papillae – rounded with taste
buds
 Circumvallate papillae – large papillae with
taste buds
 Taste buds are found on the sides of
papillae
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.49
Structure of Taste Buds
 The specific cells that respond to chemical
dissolved in saliva are epithelial cells Gustatory cells are the receptors
 Surrounded by supporting cells in the taste
bud
 Have gustatory hairs (long microvilli)
 Hairs are stimulated by chemicals dissolved in
saliva
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.50
Structure of Taste Buds
 Impulses are carried to the gustatory
complex by several cranial nerves
because taste buds are found in different
areas
 Facial nerve – anterior tongue specifically
 Glossopharyngeal nerve
 Vagus nerve
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Slide 8.51
Anatomy of Taste Buds
Figure 8.18
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Slide 8.52
Taste Sensations
 Sweet receptors
 Sugars
•May
respond
to
the
OH
 Saccharine
 Some amino acids
 Sour receptors
 Acids
 May respond to the H+
 Bitter receptors
 Alkaloids
 Salty receptors
 Metal ions in solution
Copyright © 2003 Pearson Education, Inc. publishing as Benjamin Cummings
Slide 8.53
Developmental Aspects of the
Special Senses
 Formed early in embryonic development
 Eyes are outgrowths of the brain
 All special senses are functional at birth
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Slide 8.54