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Chapter 50 An Introduction to Ecology and the Biosphere PowerPoint Lectures for Biology, Seventh Edition Neil Campbell and Jane Reece Lectures by Chris Romero Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings What you need to know! • The role of abiotic factors in the formation of biomes. • Features of freshwater and marine biomes. • Major terrestrial biomes and their characteristics. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Overview: The scope of Ecology! • Ecology – Is the scientific study of the interactions between organisms and the environment • These interactions – Determine both the distribution of organisms and their abundance Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Ecology – Study of organisms/environment (50.1) • Ecology – Has a long history as a descriptive science – Is also a rigorous experimental science • The environment of any organism includes – Abiotic, or nonliving components – Biotic, or living components – All the organisms living in the environment, the biota Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Environmental components – Affect the distribution and abundance of organisms Kangaroos/km2 > 20 10–20 5–10 1–5 0.1–1 < 0.1 Limits of distribution Figure 50.2 Climate in northern Australia is hot and wet, with seasonal drought. Southern Australia has cool, moist winters and warm, dry summers. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Red kangaroos occur in most semiarid and arid regions of the interior, where precipitation is relatively low and variable from year to year. Southeastern Australia has a wet, cool climate. Tasmania Subfields of Ecology • Organismal ecology – Studies how an organism’s structure, physiology, and (for animals) behavior meet the challenges posed by the environment Figure 50.3a (a) Organismal ecology. How do humpback whales select their calving areas? Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Population ecology – Concentrates mainly on factors that affect how many individuals of a particular species live in an area (b) Population ecology. What environmental factors affect the reproductive rate of deer mice? Figure 50.3b Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Community ecology – Deals with the whole array of interacting species in a community (c) Community ecology. What factors influence the diversity of species that make up a particular forest? Figure 50.3c Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Ecosystem ecology – Emphasizes energy flow and chemical cycling among the various biotic and abiotic components (d) Ecosystem ecology. What factors control photosynthetic productivity in a temperate grassland ecosystem? Figure 50.3d Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Landscape ecology – Deals with arrays of ecosystems and how they are arranged in a geographic region Figure 50.3e (e) Landscape ecology. To what extent do the trees lining the drainage channels in this landscape serve as corridors of dispersal for forest animals? Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • The biosphere – Is the global ecosystem, the sum of all the planet’s ecosystems Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Interactions with organisms and environments (50.2) • Ecologists – Have long recognized global and regional patterns of distribution of organisms within the biosphere – Some things can limit the distribution of specific species (like the kangaroos) Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Dispersal and Distribution • Dispersal – Is the movement of individuals away from centers of high population density or from their area of origin – Contributes to the global distribution of organisms – Affected by Biotic and Abiotic Factors Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Natural Range Expansions • Natural range expansions – Show the influence of dispersal on distribution New areas occupied Figure 50.7 Year 1996 1989 1974 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Biotic Factors • Biotic factors that affect the distribution of organisms may include – Interactions with other species – Predation – Competition Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • A specific case of an herbivore limiting distribution of a food species EXPERIMENT W. J. Fletcher tested the effects of two algae-eating animals, sea urchins and limpets, on seaweed abundance near Sydney, Australia. In areas adjacent to a control site, either the urchins, the limpets, or both were removed. RESULTS Fletcher observed a large difference in seaweed growth between areas with and without sea urchins. 100 Sea urchin Seaweed cover (%) 80 Both limpets and urchins removed Only urchins removed 60 Limpet 40 Only limpets removed Control (both urchins and limpets present) 20 Removing both limpets and urchins or removing only urchins increased seaweed cover dramatically. Almost no seaweed grew in areas where both urchins and limpets were present, or where only limpets were removed. 0 August 1982 Figure 50.8 February 1983 August 1983 February 1984 CONCLUSION Removing both limpets and urchins resulted in the greatest increase of seaweed cover, indicating that both species have some influence on seaweed distribution. But since removing only urchins greatly increased seaweed growth while removing only limpets had little effect, Fletcher concluded that sea urchins have a much greater effect than limpets in limiting seaweed distribution. Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Abiotic Factors • Abiotic factors that affect the distribution of organisms may include – Temperature – Water – Sunlight – Wind – Rocks and soil Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings 4 Abiotic Factors make up Climate Climate • Climate - Is the prevailing weather conditions in a particular area • Climate patterns can be described on two scales: – Macroclimate - patterns on the global, regional, and local level – Microclimate - very fine patterns, such as those encountered by the community of organisms underneath a fallen log Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Sunlight intensity – Plays a major part in determining the Earth’s climate patterns LALITUDINAL VARIATION IN SUNLIGHT INTENSITY North Pole 60N Low angle of incoming sunlight 30N Tropic of Cancer Sunlight directly overhead 0 (equator) Tropic of Capricorn 30S Low angle of incoming sunlight 60S South pole Figure 50.10 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Atmosphere SEASONAL VARIATION IN SUNLIGHT INTENSITY 60N June solstice: Northern Hemisphere tilts toward sun; summer begins in Northern Hemisphere; winter begins in Southern Hemisphere. 30N 0 (equator) March equinox: Equator faces sun directly; neither pole tilts toward sun; all regions on Earth experience 12 hours of daylight and 12 hours of darkness. 30S Constant tilt of 23.5 September equinox: Equator faces sun directly; neither pole tilts toward sun; all regions on Earth experience 12 hours of daylight and 12 hours of darkness. Figure 50.10 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings December solstice: Northern Hemisphere tilts away from sun; winter begins in Northern Hemisphere; summer begins in Southern Hemisphere. • Air circulation and wind patterns – Play major parts in determining the Earth’s climate patterns GLOBAL AIR CIRCULATION AND PRECIPITATION PATTERNS 60N 30N Descending dry air absorbs moisture 0 (equator) 30S 0 60S Figure 50.10 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Ascending moist air releases moisture Descending dry air absorbs moisture Arid zone Tropics Arid zone Aquatic Biomes (50.3) • Varying combinations of both biotic and abiotic factors – Determine the nature of Earth’s many biomes • Biomes – Are the major types of ecological associations that occupy broad geographic regions of land or water Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • The examination of biomes will begin with Earth’s aquatic biomes 30N Tropic of Cancer Equator Tropic of Capricorn Continental shelf 30S Key Figure 50.15 Lakes Rivers Estuaries Coral reefs Oceanic pelagic zone Intertidal zone Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Abyssal zone (below oceanic pelagic zone) • Aquatic biomes – Account for the largest part of the biosphere in terms of area – Can contain fresh or salt water • Oceans – Cover about 75% of Earth’s surface – Have an enormous impact on the biosphere Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Many aquatic biomes – Are stratified into zones or layers defined by light penetration, temperature, and depth Intertidal zone Neritic zone Littoral zone Limnetic zone 0 Oceanic zone Photic zone 200 m Continental shelf Pelagic zone Benthic zone Photic zone Aphotic zone Pelagic zone Benthic zone Aphotic zone 2,500–6,000 m Abyssal zone (deepest regions of ocean floor) (a) Zonation in a lake. The lake environment is generally classified on the basis of three physical criteria: light penetration (photic and aphotic zones), distance from shore and water depth (littoral and limnetic zones), and whether it is open water (pelagic zone) or bottom (benthic zone). Figure 50.16a, b Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings (b) Marine zonation. Like lakes, the marine environment is generally classified on the basis of light penetration (photic and aphotic zones), distance from shore and water depth (intertidal, neritic, and oceanic zones), and whether it is open water (pelagic zone) or bottom (benthic and abyssal zones). Vertical Stratification • Aquatic Biomes = vertical stratification which forms the following areas: – Photic zone – where there is enough light for photosynthesis (aphotic zone – not enough light for photosynthesis) – Benthic zone – located at bottom of biome (made up of sand, inorganic matter, and organic sediment (detritus)) – Thermoclines – narrow layers of fast temp change that separates warm upper layer with cold deeper layer Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Types of Aquatic Biomes • Freshwater – Standing bodies of water (lakes and wetlands) – Moving bodies of water (streams and rivers) • Marine Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Lakes • Communities = distributed according to waters depth – 2 Zones • Littoral zone – well lit shallow waters near shore • Limnetic zone – well-lit open surface waters farther from shore – Oligotrophic lakes – deep, nutrient poor, oxygen rich lakes – Eutrophic lakes – shallow, high nutrient content, low oxygen lakes Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Lakes LAKES Figure 50.17 An oligotrophic lake in Grand Teton, Wyoming Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings A eutrophic lake in Okavango delta, Botswana • Wetlands WETLANDS Figure 50.17 Okefenokee National Wetland Reserve in Georgia Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Streams and Rivers • Prominent physical attribute = current • Great diversity of organisms distributed in vertical zones from headwaters to mouth • Estuaries – areas where freshwater streams or rivers merge with the ocean Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Streams and rivers STREAMS AND RIVERS Figure 50.17 A headwater stream in the Great Smoky Mountains Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings The Mississippi River far form its headwaters • Estuaries ESTUARIES Figure 50.17 An estuary in a low coastal plain of Georgia Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Intertidal zones INTERTIDAL ZONES Figure 50.17 Rocky intertidal zone on the Oregon coast Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Marine Biomes • Include 2 zones and 2 Biomes: – Intertizal zones – where land meets water; periodically submerged and exposed by tides – Neritic zone – beyond intertidal zone; shallow water over continental shelves – Pelagic biome – vast realm of open blue water found past the continent shelves – Coral reef – biome created by group of cnidarians that secrete hard calcium carbonate shells; among most productive ecosystems on Earth! Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Oceanic pelagic biome OCEANIC PELAGIC BIOME Figure 50.17 Open ocean off the island of Hawaii Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Coral reefs CORAL REEFS Figure 50.17 A coral reef in the Red Sea Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Marine benthic zone MARINE BENTHIC ZONE Figure 50.17 A deep-sea hydrothermal vent community Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Climate Determines Terrestrial Biomes (50.4) • Climate – Is particularly important in determining why particular terrestrial biomes are found in certain areas Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Climate and Terrestrial Biomes • Climate has a great impact on the distribution of organisms, as seen on a climograph Temperate grassland Desert Tropical forest Annual mean temperature (ºC) 30 Temperate broadleaf forest 15 Coniferous forest 0 Arctic and alpine tundra 15 100 Figure 50.18 200 300 Annual mean precipitation (cm) Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings 400 • The distribution of major terrestrial biomes (these are the biomes for your “Biomes Chart”) 30N Tropic of Cancer Equator Tropic of Capricorn 30S Key Tropical forest Figure 50.19 Savanna Desert Chaparral Temperate grassland Temperate broadleaf forest Coniferous forest Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Tundra High mountains Polar ice Terrestrial Biomes List • Savannah • Desert • Chaparral • Temperate grassland • Temperate broadleaf forest • Coniferous Forest • Tundra • Tropical Forest Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Tropical forest TROPICAL FOREST Figure 50.20 A tropical rain forest in Borneo Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Desert DESERT Figure 50.20 The Sonoran Desert in southern Arizona Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Savanna SAVANNA Figure 50.20 A typical savanna in Kenya Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Chaparral CHAPARRAL Figure 50.20 An area of chaparral in California Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Temperate grassland TEMPERATE GRASSLAND Figure 50.20 Sheyenne National Grassland in North Dakota Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Coniferous forest CONIFEROUS FOREST Rocky Mountain National Park in Colorado Figure 50.20 Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Temperate broadleaf forest TEMPERATE BROADLEAF FOREST Figure 50.20 Great Smoky Mountains National Park in North Carolina Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings • Tundra TUNDRA Figure 50.20 Denali National Park, Alaska, in autumn Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings