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Name ____________________________________________________________ Test Date ____________________ UNIT X - EVOLUTION I. THE THEORY OF EVOLUTION (pp. 418-386) The theory of evolution is one of the most fundamental concepts in Biology. Evolution is defined as __________________ in a _______________________________________________ over time. The scientist considered as the founder of modern evolutionary theory is _____________________________________. A. History of Evolutionary Theory – During the 1700s, several scientists began challenging the idea of a world in which changes did not occur. These scientists and their hypotheses were very important to Darwin’s work. 1. Gradualism - ______________________ first suggested that the planet was much ____________________ than previously thought; began to find evidence that _________________ were slowly, but constantly taking place. 2. Malthus – Published an essay that had a huge impact on Darwin. Proposed that organisms _________- reproduce; in other words, reproduce at a ______________________ rate than resources can supply. 3. Lamarck Lamarck was one of the first scientists to propose a mechanism for evolution; that is, the _____________________ in a _________________________ of organisms over ____________. His major hypotheses included: Tendency Toward Perfection – Stated that organisms were continually changing in order to _____________________________________. Use and Disuse – Changes in __________________ and/or __________________ of a structure in an organism was a response to use or disuse. Structures used extensively ___________________________ and structures used less frequently ________________________________________________. Inheritance of _________________ Traits. Ex: Giraffes with long necks B. Charles Darwin (1809 – 1882) 1. Darwin’s History Darwin’s data was collected on a 5-year journey around the world on the HMS ________________. He made observations and collected data throughout the journey. He used this data to propose a ______________________ to explain the diversity he saw. The area that had the greatest impact on Darwin was the _____________________ Islands due to the differences he saw in the same animals living on different islands. 2. Darwin’s Theory of Natural Selection 4 Principles: Variation - Members of a population often vary greatly in their ____________________________. Heritability - Traits are inherited from _________________________ to _____________________________. Overproduction - All species are capable of producing ____________________ offspring than _________________________________. Due to a lack of _________________ or other resources, many of the offspring ________________________. Reproductive Advantage - Organisms with favorable ____________________________ tend to survive and ____________________________; thereby leaving more descendants than other individuals. This will result in an accumulation of these traits in the __________________________________, changing the original make-up of the population. 3. Darwin’s Theory of Descent with Modification Each living species has descended (with changes) from other species over time. Implies that all living things are related to one another. Also known as common descent. According to this principle, there would be a single “tree of life” for all living things. This tree of life is commonly known as a _______________________________________. These can also be referred to as cladograms. 4. Darwin’s Legacy Did not publish his findings for years Alfred Wallace – formed identical _______________________ based on his research. Sent his manuscript to Darwin, and finally Darwin was persuaded to publish his own conclusions Released “_________________________________”, still considered one of the greatest scientific studies ever II. THE PRINCIPLES OF EVOLUTIONARY THEORY (pp. 437-438) A. Evolution occurs because of natural selection - a mechanism for change that occurs when organisms with __________________________ characteristics for a particular environment __________________________________, ____________________________, and pass these characteristics on to ____________________________________. B. The ability of an organism to survive and reproduce in its environment is known as ____________________________. C. Fitness is based on _______________________________. An adaptation is any trait that aids in the __________________________ and _____________________________ of an organism. Examples of adaptations are _______________________________________________________________________________________________ D. As organisms __________________ and adapt, ______________________ may occur. Speciation is the formation of new species - a group of similar organisms that _____________________________ with one another and produce _______________________________________ that is, babies that can make babies! Isolating Mechanisms allow for the gene pools to become ________________ so they can form a new ____________. Reproductive Isolation-as a new _________________________evolves, population become ____________________ isolated from each other. Behavioral Isolation-Population is capable of _____________________________________, but have different ______________________________ rituals. EX: Eastern and Western Meadowlark Geographic Isolation-_____________ populations are separated by geographic barriers and evolve where they can no longer ________________________________. EX: Abert and Kaibab squirrels of the Grand Canyon Temporal Isolation-______________________________ reproduce at different times even though capable. EX: Three orchids of the rain forest, release pollen only 1 day and not the same day E. The failure of an organism to ___________________________ to changes in its environment will ultimately lead to its ________________________ because of __________________________________________. III. EVIDENCE FOR EVOLUTION (pp. 423-429) A. Fossils – Fossils are _________________________________________________________________ Fossils provide a record of earlier life and evidence that evolution has occurred. B. Biogeography – Variations are seen in the same types of animals based on their ___________________________. In addition, there are some organisms that live in very different locations but they have similar characteristics because _____________________________________________, for example, climate, food sources, etc. C. Homologous Structures – Scientists use anatomical studies of different organisms for evidence of evolutionary relationships. For example, appendages that are very similar in structure, but differ in function are known as ___________________________ structures. Examples of homologous structures are __________________________________________________________________________________________. D. Vestigial Structures - A structure that is reduced in function in a living organism, but may have been used by an ancestor is known as a _______________________ structure. A structure may become vestigial when an organism changes in form or behavior. Examples are __________________________________________________________. E. Embryology – Similarities in the structures of developing ___________________________ of different organisms are considered to be proof of a close evolutionary relationship. F. DNA Sequencing – Scientists use DNA studies to determine the evolutionary relationship between organisms. The more similar the DNA, __________________________________________________ IV. MECHANISMS FOR EVOLUTION (pp. 431-439) Evolution does not occur in an individual; instead it refers to ________________ that occur in a ____________________ over time. There must be mechanisms available for _______________________ changes to occur: A. Mutation – A mutation is a _____________________________________. Although mutations are most often _________________________, sometimes the resulting change in _____________________________ may be beneficial to an organism under certain conditions. If the change occurs in the ____________________, this change will be passed onto the offspring of that organism. A positive mutation that provides a survival advantage is known as an __________________________. B. Recombination – Leads to increased genetic variation as a result of ______________________________ during ______________________________ of ____________________________. C. Gene Flow – Gene flow occurs when organisms from one community migrate to another. This introduces new __________________________ into the ____________________________ which can lead to a change in the genetic make-up of the population. D. Genetic Drift – This describes a situation in which change in a population is magnified because the population size is very __________________________________. Causes of genetic drift include: Bottleneck effect – large portion of population ______________________________________________ Founder effect – segment of population moves to new ________________________ V. MACROEVOLUTION (pp. 439-440) Refers to large-scale evolutionary pattern and processes that occur over long periods of time. A. Extinction-More than _______% of all species that have ever lived are now ____________________, which means the species has died out. Darwin proposed possible reasons with competition for ___________________and environmental change. ________________extinction has occurred several times, wiping out entire __________________________. _____________________, volcanic action, and __________________________movement has been blamed. Mass extinctions clear the way for _______________________ of other species. B. Adaptive radiation-the process where a ____________________species has evolved through _________________________ into diverse forms that live in _________________________ways. EX: ____________ ______________________________________________________________ C.____________________ _____________________can have an evolutionary “side effect”, producing unrelated species that ________________ similar to one another. This process is known as ____________________________________ and occurs in ________________________ and animals. These organisms have _____________________________ structures. EX:_____________________________________________ D. _________________________ Equilibrium is another pattern of evolution. Unlike ___________________________ punctuated equilibrium is characterized with long periods of _________________________ interrupted by brief periods of ______________________change. It is controversial but known that evolution does occur at different rates. VI. Modern Evolutionary Classification (pp. 440-441) Linnaeus tried to group organisms according to biologically important characteristics, but his system had limitations and problems. A. Darwin’s ideas about descent with modification gave rise to the study of _______________________________, or evolutionary relationships among organisms. Biologists now group organisms into categories that represent lines of _______________________________, or phylogeny, not just physical similarities. B. Cladistic Analysis-identifies and considers only those characteristics of organisms that are _____________________________________________ -new characteristics that arise as lineages evolve over time. These recent characteristics are called ______________________________________________. C. Derived characters can be used to construct a ______________________________________, a diagram that shows the evolutionary relationships among a group of organisms. Cladograms-are useful tools that help scientists understand how one lineage ____________________________ from another in the course of evolution. Think of them as a family tree showing __________________________________relationships among a group of organisms. D. Similarities in Genetic Material ____________ and _____________ are so similar across all forms of life, these molecules provide an excellent way of comparing organisms at their most basic level_____________________________________________. _________________________________________ in DNA can be used to help determine classification and _______________________________________relationships. DNA___________________________________ using ________________________________________ is a technique that can be used to analyze DNA and determine these evolutionary relationships. ________________________________________________ or reading of the DNA of organisms allow for comparison of the DNA of different organisms to trace the history of genes over ________________ of years. VII. THE EVOLUTION OF THE CELL – THE ENDOSYMBIOTIC THEORY (pp. 406-407)