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Amazing Arthropods! The Little Creatures Who Rule the World General Characteristics • • • • • Segmented bodies Jointed appendages Exoskeleton Highly cephalized Open circulatory system Exoskeleton - advantages • Enables great strength (internal muscle attachment). • Waterproof - enables life on land. • Protective armor. Exoskeleton - limitations • Must be molted (shed) in order to grow. Recently molted animals are vulnerable. • http://www.youtube.co m/watch?v=mF6NgM BcNCM • Heavy - limits overall size. Cephalization • Specialized mouthparts sensory, food-handling. • Antennae - sense chemicals, vibrations. • Compound eyes - many individual units, each w/own lens, good at detecting motion, color. • Simple eyes - detect light intensity. Arthropods: Class Crustacea • Crabs, lobster, shrimp, barnacles, etc. • Mostly aquatic • Two body divisions, cephalothorax & abdomen • http://www.youtube.c om/watch?v=nmfYW 7wAbQM • Most crustaceans are scavengers or predators. • Chelipeds – strong claws for grasping & cutting or tearing • http://www.youtube.com/watch?v=b Sd_5R04f9Q • http://www.youtube.com/watch?v=m VuMousxF70 • 4 prs. Walking legs • Swimmerets – on abdomen, move water over gills & or eggs. Crayfish Anatomy Internal Anatomy & Senses • • • • • Green glands – excretory organs in head Open circulatory system Gills – attached to each walking leg, moves H2O over them as it walks. Ventral nerve cord Compound eyes – many lenses working together, good for detecting movement. • Statocysts – detect orientation (up & down) as gravity pulls on grains of calcium carbonate inside. • Males have specialized swimmerets for sperm transfer – internal fertilization. • Antennae – touch, chemicals Arthropods: Class Arachnida • Spiders, scorpions, mites, ticks • Mostly terrestrial • Two body divisions, cephalothorax & abdomen • 4 prs. Walking legs • Carnivorous or parasitic Spider Anatomy • Book lungs for respiration – large surface area. • Spinnerets – silk glands produce silk as liquid, emerges from spinnerets & solidifies on contact with air. • Spider silk is strongest natural substance known. • Malpighian tubules – excretory organs, extensions of gut. • Chelicerae house fangs to inject venom. • Stronger than steel • Many uses – webs, parachutes, guidelines, camouflage • http://www.youtube.com/watch?v=y SXCdcnKBgg • http://www.youtube.com/watch?v=i BjjNaRzdEk&NR=1&feature=fvwp Spider Webs & Silk • Millipedes • 2 prs. legs/body segment • Herbivorous Arthropods: Class Diplopoda • • • • Centipedes 1 pr. Legs/segment Carnivorous Venomous Arthropods: Class Chilopoda Arthropods: Class Insecta • • • • General Characteristics 3 body divisions - head, thorax, abdomen. 1 pr. antennae 3 prs. Legs on thorax, 1 pr./segment. 2 pr. wings (usually), 1 pr./1st two thoracic segments. Insect Adaptations for Success Flight Easier to: • disperse to new areas. • find food. • find mates. • escape predators. Insect Adaptations for Success Small Size • Individuals need fewer resources. • Many individuals can inhabit small area without competition. • Easier to hide. Insect Adaptations for Success • • • • Exoskeleton Great muscular leverage. Waterproof. Protective. Flexible at joints. Insect Adaptations for Success Highly adaptable • short lifespan & generation time, many offspring. • Frequent mutations, some beneficial, so species can respond to environmental changes. Insect Digestive System Specialized Mouthparts • chewing – beetle, grasshopper • piercing/sucking – mosquito, true bugs • sponging - house fly • siphoning - butterfly Insect Digestive/Excretory System • Crop - storage • Gizzard - grinds food • midgut (stomach) digestion. • Gastric cecae - digestive glands. • Hindgut - nutrient absorption • Malpighian tubules removes waste from blood, deposits in rectum. • Open • Dorsal aorta • Heart posterior, pumps blood forward. • Blood empties into coelom, bathes tissues as it flows to posterior. Insect Circulatory System Insect Respiratory System • Air intake through spiracles on sides of abdomen. • Air pulled into system of tubes (tracheae) that transport air directly to tissues. • Oxygen diffuses into tissues, CO2 out. Nervous System & Sense Organs • Nerve ganglia - “brain” • Ventral Nerve cord. • Tympanum - sound sensing organ on abdomen or foreleg - not all insects have them. Reproductive System • Sperm deposited into female’s seminal receptacle. • Eggs fertilized internally. • Ovipositor used to create hole in substrate and/or deposit eggs. Metamorphosis Incomplete • 3 stages – egg, nymph, adult • Nymph – immature form, resembles adult, but w/ underdeveloped sexual organs. • Ex.- dragonflies, grasshoppers, true bugs, hoppers (Homoptera) Complete Metamorphosis • 4 stages – egg, larva, pupa, adult, each distinctly different. • Larva – specialized for feeding & growth. • Pupa – transformation stage – appears inactive, but big changes internally. • Adult – specialized for dispersal, mating, & feeding – different food than larva, no competition. • Butterflies, bees, beetles, flies. Importance of Metamorphosis • One main factor in insect success. • Can exploit seasonally-available foods. • Different stages requiring lots of energy are separate - no competition between adults & larvae for food & space. • Insects can survive harsh weather as eggs or pupae. • Aggression – stings or chemicals. • Camouflage or hiding. • Advertise danger Mullerian mimicry – all dangerous insects have similar coloration – predators only need to learn once. Batesian mimicry – harmless insect evolves to resemble dangerous one. (monarch/viceroy) Defense Other Forms of Mimicry • Looking like part of the environment (stick, leaf, thorn, etc.) helps insects escape notice. Communication • Social insects have the most elaborate communication. • Communication may be by scent (pheromones), sound, touch, or light (fireflies). Social Behavior • Includes ants, termites, and colonial bees and wasps • Caste system – division of labor • Behavior is instinctive, not learned • Workers – responsible for all tasks of the colony except reproduction. • Drones – males, mate w/ queen (from another colony) • Queen – female, reproduction only. Honeybee Castes Workers • Live about 6 wks. • Tasks change w/ age. • 1st task – “nurse” bees, feed larvae, drones, & queen. • Queen & youngest larvae get royal jelly. • 2nd task – start producing wax to build & repair honeycomb. Also remove waste & dead bees, guard hive, & circulate air in hive. • Last task – gather nectar & pollen Queen • Lives up to 5 yrs. • Queen develops from larva fed a continuous diet of royal jelly. • Adult queen secretes “queen factor”, pheromone preventing other female larvae from sexually maturing. • Mates once in her life on mating flight, stores sperm to fertilize eggs for rest of life. • Overcrowded hive, old queen takes half the workers & swarms to start new colony. Worker left behind produce more queens that fight for dominance. Drones • Males, develop from unfertilized eggs. • Large eyes – the better to see you with, my queen! • Contribute nothing to hive except reproduction. • Tolerated and fed by workers during seasons of plenty, but killed or driven from hive when food is scarce (winter). • Only a few hundred drones, compared to several thousand workers. Waggle Dance & Round Dance • < of straight run to vertical equals < between food source to sun. • # waggles in straight run indicates distance. • Round dance indicates food nearby, but not direction. Major Orders • Orthoptera “straight-wing” • Grasshoppers, crickets • Strong, muscular hind legs for jumping Major Orders • Odonata – “toothed wing” • Dragonflies, Damselflies • Long, narrow, membranous wings Major Orders • • • • Hemiptera – “Half-wing” True bugs Half-membranous, half-leathery forewings Large triangle on back Major Orders • Homoptera – “Like-wing, same-wing” • Cicadas, aphids, tree hoppers, frog hoppers • Hold wings at angle over back to form “roof” Major Orders • Coleoptera – “sheath wing” – beetles • Largest order, great diversity • Forewings hard or leathery, cover most or all of abdomen, meet in straight line on back Major Orders • Diptera – “two-wing” • Flies, gnats, mosquitoes • One pair of wings Major Orders • Hymenoptera – “Membrane-wing” • Bees, ants, wasps • Skinny “waist” Major Orders • Lepidoptera – “scale wing” • Butterflies, moths • Wings covered in scales