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Lecture -8: Ecological Succession ENV 107: Introduction to Environmental Science Dr. A.K.M. Saiful Islam ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Case Study: Restoring Abandoned Mine Lands in Great Britain In Great Britain, approx. 55,000 ha have been damaged by mining. Recently, in one damaged area (British Peak District National park) programs have been initiated to restore these damaged land. First attempt to restore this area used a modern agricultural approach: heavy application of fertilizers and planting of fast growing agriculture grasses. These grasses green on the good soil of a level firm field. But after a short period of growth, the grasses died! ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Lesson learned On the poor soil, leached of its nutrients and lacking organic matter, erosion continued and the fertilizers that had been added were soon leached away by water runoff. Ecological succession, the natural process of establishment or reestablishment of an ecosystem. Instead of planting fast growing but vulnerable agricultural grasses, ecologist planted slow-growing native grasses that were known to be adapted to minerally deficient soils and to the harsh conditions that exist in cleared areas. ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Ecological Succession Recovery of ecosystem occurs naturally through a process called ecological succession. The natural recovery can occur if the damage is not too great. Sometimes the rate of recovery is long in comparison to human desired. The disturbance of nature not always usually by humans. Natural disturbances included: storms, fires have always been a part of the environment. There have been 2 kinds of succession: Primary succession: is initial establishment and development of an ecosystem. Secondary succession: is reestablishment of an ecosystem. ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Examples of Primary succession Forests that develop on new lava flows. Ferns are among the first plants to reestablish after a new volcanic flow. (a) Re-vegetation on new lava flows on Hawaii. (b) Re-vegetation of the land at the retreating edge of a New Zealand glacier ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Examples of Secondary Succession A forest that developed on an abandon pasture or one grows after a hurricane, flood or fire. (a) Poconos pasture abandoned filed (c) Mature forests developed ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Stages of Succession Early successional species: In forest areas, plants that are rapid growing and short lived Late successional species: Plant species that dominate later stages of succession tend to be slower growing and longer lived. These plants do comparatively ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Graph of changes in biomass and diversity with succession ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Graph of changes in biomass and diversity with succession (Contd..) ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Patterns of interaction There are at least three patterns of interactions among earlier and later species in succession. 1. Facilitation 2. Interference 3. Life History Differences There is actually a fourth possibility: succession never occurs or chronic patchiness. 4. Chronic Patchiness ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam 1. Facilitations One species can prepare the way for the next (and many even be necessary for the occurrence of the next). Pine provide shade and act as “nurse trees” for oak Pines do well in openings. If there is no pine few or no oaks will grow. The pine facilitate the entrance of oak ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam 2. Interference Early successional species may in some way prevent the entrance of later successional species. Some grasses that grows in open areas form dense Mats that prevent seeds of trees from reaching the soil ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam 3. Life History Differences One species may not affect the time of entrance of another; two species may appear at different times during succession. ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam 4. Chronic patchiness Succession never occurs and the species that enters first remains until the next disturbance. The null condition, neither positive nor negative interactions ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Some special cases of Ecological Succession 1. 2. 3. 4. Succession among the ocean tides Wetland succession Bog Succession Sand Dune Succession ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam 1. Succession among the ocean tides In the ocean, succession occurs where is relation constancy of the environment such as between the high and low tidemarks along a rocky shore or in a coral reef. Where the ocean environment is constantly changing, succession does not appear to occur. For example, there is no perceptible succession in the middle of the ocean, where open ocean waters are continually stirred by winds, waves, and currents. ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam 2. Wetland succession From a geologic point of view, a pond is a temporary feature of a landscape, eventually filling in with sediments. Ponds are common in areas subject to large-scale geological disturbances, such as glaciation, that shift the drainage patterns of streams and create depressions and dams. This is why Minnesota, which lies within the heavily glaciated area of North America, is called the Land of 10,000 lakes. ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam 3. Bog Succession A bog is a body of water with acid waters and little if any surface out-flow, so that the waters have little current. Often, a bog has sphagnum moss and inlets but no surface out-lets. Succession in a bog is a process that begins with open water and ends with a forest . ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Bog Succession ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam 4. Sand Dune Succession Plant succession on sand dunes along beaches is a worldwide phenomenon, important along the shores of Great Lakes in North America, in cape Cod, Massachusetts, where Henry David Thoreau observed it in the nine-teeth century, along the coasts of Australia, as well as in the famous Donana National Park in southern Shores of Spain. Dunes along the shores of Lake Michigan were among the first sites where succession was studied early in the twentieth century. ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Dune succession on the shore of Lake Michigan ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Succession and Chemical Cycling On the land, there is generally an increase in the storage of chemical elements (Nitrogen, phosphorus, calcium essential for plant growth) during the progression from the earliest stages of succession to middle or late succession. ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam Change in Chemical Cycling during Disturbance When an ecosystem is disturbed by fire, storms, or human actions, changes occur in chemical cycling. For example, when a forest is burned, complex organic compounds such as wood are converted to smaller inorganic compounds. ENV 107:Introduction to Environmental Science © Dr. Akm Saiful Islam How can be Evaluate Constructed Ecosystem 1. Establishment of tide channels with sufficient fish to serve as food for the California least tern. 2. Establishment of a stable or increasing population of salt-marsh bird’s beak for three years. 3. Establishment of habitat for the lightfooted successful foraging and nesting of at least two breeding pairs.