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Chapter 52 An Introduction to Ecology and the Biosphere PowerPoint® Lecture Presentations for Biology Eighth Edition Neil Campbell and Jane Reece Lectures by Chris Romero, updated by Erin Barley with contributions from Joan Sharp Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Key concepts 1. Ecology is to study “interaction” within, between, and across species, as well as the environment. 2. Ecology is an integrated discipline. Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Overview: The Scope of Ecology • Ecology is the scientific study of the interactions between organisms and the environment • These interactions determine distribution of organisms and their abundance • Ecology reveals the richness of the biosphere Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Fig. 52-1 Why do gray whales migrate? • Organismal ecology studies how an organism‟s structure, physiology, and (for animals) behavior meet environmental challenges Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • A population is a group of individuals of the same species living in an area • Population ecology focuses on factors affecting how many individuals of a species live in an area Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • A community is a group of populations of different species in an area • Community ecology deals with the whole array of interacting species in a community Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • An ecosystem is the community of organisms in an area and the physical factors with which they interact • Ecosystem ecology emphasizes energy flow and chemical cycling among the various biotic and abiotic components Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • A landscape is a mosaic of connected ecosystems • Landscape ecology deals with arrays of ecosystems and how they are arranged in a geographic region Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings • The biosphere is the global ecosystem, the sum of all the planet‟s ecosystems • Global ecology examines the influence of energy and materials on organisms across the biosphere Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Fig. 52-2 Organismal ecology Population ecology Community ecology Ecosystem ecology Landscape ecology Global ecology Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Fig. 52-3 Ecology has a long history as a descriptive science, but it is also a rigorous experimental science Trough Pipe “Dry” “Wet” “Ambient” Studying how a forest responds to altered precipitation Ecology and Environmental Issues • Ecology – Provides the scientific understanding underlying environmental issues • Rachel Carson Silent Spring (1962) “The „control of nature‟ is a phrase conceived in arrogance, born of the Neanderthal age of biology and philosophy, when it was supposed that nature exists for the convenience of man.” Copyright © 2005 Pearson Education, Inc. publishing as Benjamin Cummings Concept 52.2: Interactions between organisms and the environment limit the distribution of species • Ecologists recognize two kinds of factors that determine distribution: biotic, or living factors, and abiotic, or nonliving factors Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Fig. 52-5 Kangaroos/km2 0–0.1 0.1–1 1–5 5–10 10–20 > 20 Limits of distribution Distribution and abundance of the red kangaroo in Australia, based on aerial surveys Fig. 52-6 Flowchart of factors limiting geographic distribution Why is species X absent from an area? Yes Does dispersal limit its distribution? No Area inaccessible or insufficient time Does behavior limit its distribution? Yes Habitat selection Yes No Do biotic factors (other species) limit its distribution? No Predation, parasitism, Chemical competition, disease factors Do abiotic factors limit its distribution? Water Oxygen Salinity pH Soil nutrients, etc. Temperature Physical Light factors Soil structure Fire Moisture, etc. Fig. 52-7 Natural range expansions show the influence of dispersal on distribution Current 1970 1966 1965 1960 Dispersal of the cattle egret in the Americas 1961 1943 1958 1951 1956 1970 1937 Species Transplants • Species transplants include organisms that are intentionally or accidentally relocated from their original distribution Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Behavior and Habitat Selection • Some organisms do not occupy all of their potential range • Species distribution may be limited by habitat selection behavior Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Fig. 52-8 Does feeding by sea urchins limit seaweed distribution? RESULTS 100 Seaweed cover (%) 80 Both limpets and urchins removed Sea urchin Only urchins removed 60 Limpet 40 Only limpets removed Control (both urchins and limpets present) 20 0 August 1982 February 1983 August 1983 February 1984 Biotic Factors • Biotic factors that affect the distribution of organisms may include: – Interactions with other species – Predation – Competition Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Abiotic Factors • Abiotic factors affecting distribution of organisms include: – Temperature – Water – Sunlight – Wind – Rocks and soil • Most abiotic factors vary in space and time Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Fig. 52-9 Fig. 52-10a Latitudinal Variation in Sunlight Intensity 90ºN (North Pole) 60ºN Low angle of incoming sunlight 30ºN 23.5ºN (Tropic of Cancer) Sun directly overhead at equinoxes 0º (equator) 23.5ºS (Tropic of Capricorn) 30ºS Low angle of incoming sunlight 60ºS 90ºS (South Pole) Atmosphere Seasonal Variation in Sunlight Intensity 60ºN 30ºN March equinox 0º (equator) June solstice 30ºS Constant tilt of 23.5º September equinox December solstice Fig. 52-10d Global Air Circulation and Precipitation Patterns 60ºN 30ºN Descending dry air absorbs moisture 0º (equator) 30ºS 60ºS 30º 23.5º Arid zone Ascending moist air releases moisture 0º Tropics Descending dry air absorbs moisture 23.5º 30º Arid zone Global Wind Patterns 66.5ºN (Arctic Circle) 60ºN Westerlies 30ºN Northeast trades Doldrums Southeast trades 0º (equator) 30ºS Westerlies 60ºS 66.5ºS (Antarctic Circle) Fig. 52-11 The great ocean conveyor belt Labrador current Gulf stream Equator Cold water Fig. 52-12 2 Air cools at 3 Cooler high elevation. air sinks over water. 1 Warm air over land rises. 4 Cool air over water moves inland, replacing rising warm air over land. Moderating effects of a large body of water on climate Fig. 52-13 How mountains affect rainfall Wind direction Leeward side of mountain Mountain range Ocean Current range and predicted range for the American beech (Fagus grandifolia) under two scenarios of climate change Current range Predicted range Overlap (a) 4.5ºC warming over next century (b) 6.5ºC warming over next century Concept 52.3: Aquatic biomes are diverse and dynamic systems that cover most of Earth • Biomes are the major ecological associations that occupy broad geographic regions of land or water • Varying combinations of biotic and abiotic factors determine the nature of biomes Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings Fig. 52-15 The distribution of major aquatic biomes 30ºN Tropic of Cancer Equator Tropic of Capricorn 30ºS Lakes Coral reefs Rivers Oceanic pelagic and benthic zones Estuaries Intertidal zones Fig. 52-16 Zonation in aquatic environments Intertidal zone Oceanic zone Neritic zone Littoral zone Limnetic zone 0 Photic zone 200 m Continental shelf Benthic zone Photic zone Benthic zone Pelagic zone Pelagic zone Aphotic zone 2,000–6,000 m Abyssal zone (a) Zonation in a lake (b) Marine zonation Aphotic zone Fig. 52-17-5 Seasonal turnover in lakes with winter ice cover Winter Summer Spring 2º 4º 4º 4º 4ºC 0º 4º 4º 4º 4º 4º Autumn 20º 18º 8º 6º 5º 4ºC 4ºC Thermocline 22º 4º 4º 4º 4º 4ºC 4º Fig. 52-18a An oligotrophic lake in Grand Teton National Park, Wyoming Fig. 52-18b A eutrophic lake in the Okavango Delta, Botswana Fig. 52-18c Okefenokee National Wetland Reserve in Georgia Fig. 52-18d A headwater stream in the Great Smoky Mountains Fig. 52-18e The Mississippi River far from its headwaters Fig. 52-18f An estuary in a low coastal plain of Georgia Fig. 52-18g Rocky intertidal zone on the Oregon coast Fig. 52-18h Open ocean off the island of Hawaii Fig. 52-18i A coral reef in the Red Sea Fig. 52-18j A deep-sea hydrothermal vent community Fig. 52-19 The distribution of major terrestrial biomes Tropical forest Savanna Desert 30ºN Tropic of Cancer Equator Tropic of Capricorn 30ºS Chaparral Temperate grassland Temperate broadleaf forest Northern coniferous forest Tundra High mountains Polar ice Fig. 52-20 Climograph Annual mean temperature (ºC) Desert Temperate grassland Tropical forest 30 Temperate broadleaf forest 15 Northern coniferous forest 0 Arctic and alpine tundra –15 0 100 200 400 300 Annual mean precipitation (cm) Fig. 52-21a A tropical rain forest in Borneo Fig. 52-21b A desert in the southwestern United States Fig. 52-21c A savanna in Kenya Fig. 52-21d An area of chaparral in California Fig. 52-21e Sheyenne National Grassland in North Dakota Fig. 52-21f Rocky Mountain National Park in Colorado northern coniferous forest Fig. 52-21g Great Smoky Mountains National Park in North Carolina temperate broadleaf forest Fig. 52-21h Denali National Park, Alaska, in autumn tundra You should now be able to: 1. Distinguish among the following types of ecology: organismal, population, community, ecosystem, and landscape 2. Explain how dispersal may contribute to a species‟ distribution 3. Distinguish between the following pairs of terms: potential and actual range, biotic and abiotic factors, macroclimate and microclimate patterns Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings 4. Explain how a body of water or mountain range might affect regional climatic conditions 5. Define the following terms: photic zone, aphotic zone, benthic zone, abyssal zone, thermal stratification, thermocline, seasonal turnover, climograph, disturbance 6. List and describe the characteristics of the major aquatic biomes Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings 7. List and describe the characteristics of the major terrestrial biomes 8. Compare the vertical layering of a forest and grassland Copyright © 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings