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Hominid Evolution Crystal A. Brandon Evolutionary Relationship Amongst Hominid Species Emergence of Hominid Species Hominid Brain Evolution 1400 1000 800 600 400 200 A fa re ns is A fr ic an us G ar hi H ab ili s E rg as te r E re ct us Sa pi en s 0 Pa n Brain Volume 1200 Estimation of Values in Figure 3.4 of text Important Brain Changes Across Hominid Species Increase in brain volume across species of Australopithecines Substantial changes in facial structure of Homo Expansion of frontal and parietal lobes of H. habilis Continued expansion from emergence of H. erectus to modern day humans Encephalization Quotient An index of brain size relative to that of an average mammal of the same body weight Ex. Chimps = 2.0 Humans = 5.0 to 6.0, on average Adaptation & Selection Pressures: Climatic Pressures Turnover Pulse Hypothesis (Vrba, 1995) Hypothesis: Large scale climatic changes result in habitat and ecological changes, resulting in pulses of speciation and extinction Adaptation & Selection Pressures: Ecological Pressures Focus on the ability of hominids to extract biological resources from the ecology improving survival prospects and support population increases and expansion Superpredator: greater ability to capture and process Patterns of migration & mass extinctions of species Evidence in nonhumans species: Foraging and complex predatory demands correlated with larger brain volume and higher EQ Adaptation & Selection Pressures: Ecological Pressures Kaplan et al. (2000): Hypothesis If the ability to extract and process biological resources was the driving force in the evolution of brain and cognition, then improvements in the ability of hominids to extract these resources should corresponds to changes in brain volume and EQ during hominid evolution. Ex. Fire and sophisticated tool use Adaptation & Selection Pressures: Social Pressures Ecological Dominance The ability to very efficiently extract biological resources from the ecology and manipulate the ecology in ways that: 1. reduce mortality risks 2. support subsequent population expansions Alexander (1989): Ecological pressures were more prominent earlier in hominid evolution, and social pressures more prominent later in hominid evolution Social Complexity & Ecological Dominance Ecological Dominance: Control of Survival-Related Resources Supporting Traits Brain and Cognition Folk Biology Folk Physics Sociality and Behavior Hunting/Foraging/Ecological Manipulation Co-evolving Traits: Function to Adapt Supporting Traits to Local Ecology Increased Parental Investment Increased Developmental Activity Social Complexity & Ecological Dominance Increase in Paternal Investment Increase in length of developmental period Increased Developmental Activity More opportunities to practice hunting and foraging Social Dynamics & Ecological Dominance Social Dynamics: Social Struggle for Control of Survival and Reproduction-Related Resources Supporting Traits Brain and Cognition Sociality and Behavior Kinship Networks Folk Psychology Reciprocal Relationships Co-evolving Traits: Function to Adapt Supporting Traits to Local Ecology Increased Parental Investment Increased Developmental Activity Social Dynamics & Ecological Dominance Kinship groups in competition with other kinship groups over resources and reproduction Kinship groups facilitate: 1. Peer relationships 2. Rough-and-tumble play Forms of Social Conflict and Competition Intrasexual Competition Male-Male Competition Female-Female Competition Within Group and Between Group Competition for Control of Resources and Position Within group competition often involving subtle manipulation of other members to gain access to resources Intersexual choice Male Choice Men will be selective in their choice of a mate because of paternal investment and paternal certainty Female Choice Women will select their mate based on male social status and male paternal competency Family Relationships Spousal Parent-offspring and sibling Conflict over maternal vs. paternal investment; resource control; and marital fidelity Extended developmental period results in conflict over parental allocation of resources Motivation to Control Human motivation to control focuses on attempts to control: 1. Social Relationships: Benefits? - protection, territorial gain, shared access to resources 2. Other People: Benefits? - reproductive potential, social power, access to resources 3. Biological and Physical Resources: Benefits? - healthier, inherited strategies to obtain resources Evolutionary Mechanisms to Support Motivation to Control Conscious-psychological mechanisms Affective mechanisms Cognitive mechanisms