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Invertebrate Chordates CHORDATES ARE ANIMALS THAT ARE CHARACTERIZED BY A NOTOCHORD, A HOLLOW DORSAL NERVE CORD, AND PHARYNGEAL THROAT SLITS. Notochord: y A long flexible supporting rod that runs through at least part of the body – usually along the dorsal surface just beneath the nerve cord. Present usually early in development to be replaced by a backbone in vertebrates. Hollow Dorsal Nerve Cord: y Runs along the dorsal surface just above the notochord (in most invertebrates nerve cords run along the ventral surface). Nerves leave this cord at regular intervals to connect with organs, muscles, and sense organs. The front end of the cord, in most chordates, develops into a brain. Pharyngeal slits: y Paired structures in the throat region of the body. In aquatic chordates – lancets and fishes – these are gill slits. Many invertebrates have gills of some sort in various places whereas terrestrial chordates that use lungs for respiration, have pharyngeal slits for a brief period of time during the development of the embryo. Tunicates: y Small marine chordates that eat plankton that they filter from the water. Only the tadpole-shaped larvae of tunicates have a notochord and a dorsal nerve cord that leave when the animal undergoes metamorphosis to become sessile, attach to the floor, and live in colonies. These animals filter feed and breathe at the same time through a pharyngeal basket pierced by gill slits. Lancelets: y Live in the sandy body of shallow tropical oceans. They are fishlike with a definite head. y Feeding – pass water through their pharynx where food gets stuck in mucus. This mucus is then swallowed into the digestive tract that passes straight to the anus near the tail. Lancelets have no jaw. y Respiration – they have a mouth that opens to a long pharyngeal region with up to 100 pairs of gill slits. Lancelets Con’t: y Vascular system – Simple primitive heart that pumps blood through vessels in a closed circulatory system. Show evidence of segmentation as their muscles are organized into V – shaped units that are on either side of the body. Each unit receives a branch from the main nerve cord. This segmented nerve and muscle organization is found in all living vertebrates. y Movement – they have no appendages and move by bending their bodies back and forth.