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FOSS Ecoscenarios: El Yunque National Forest General Information Page 1 of 7 INTRODUCTION Biome: Tropical Rain Forest In the northern region of the Caribbean Sea, 1,200 kilometers (km) southeast of Florida, lies the island of Puerto Rico. Nearly 60 percent of the island is covered with mountains. It is also home to many different ecosystems including hot deserts, mangrove forests, grasslands, freshwater rivers, coral reefs, caves, and tropical rain forests. The Sierra de Luquillo Mountains are clustered in the northeastern part of Puerto Rico. Tropical storms build as they come across the Atlantic Ocean and drop large amounts of rainfall on the Sierra de Luquillo. The result is a lush tropical rain forest ecosystem. The tropical rain forest of the Sierra de Luquillo was first protected in 1876, when Puerto Rico was under Spanish rule. By declaring it a reserve, the Spanish protected the trees so that enemies could not use their lumber to make ships that might fight against them. It was one of the first reserves in the Western Hemisphere. Courtesy of Father Alejandro Sánchez The canopy of trees in El Yunque forms a green carpet that covers the mountains, Sierra de Luquillo. In 1898, Puerto Rico became a self-governing territory of the United States. Eventually the eastern preserve became part of the US Forest Service. It was ultimately renamed El Yunque National Forest in honor of the native Tiano people, who called the cloud-shrouded FOSS Populations and Ecosystems Course, Second Edition © The Regents of the University of California Can be duplicated for classroom or workshop use. Version date 12/2015 FOSS Ecoscenarios: El Yunque National Forest General Information Page 2 of 7 mountaintops “yuke,” meaning “white lands.” It is also the name of the tallest mountain in the Sierra de Luquillo range. El Yunque National Forest covers approximately 113 square kilometers (km2) of mountainous tropical rain forest. El Yunque offers many ecosystem services. Worldwide, tropical rain forests cover 2 percent of Earth’s surface; yet contain 50 percent of all plant and animal species. El Yunque is the only tropical rain forest managed by the US Forest Service and is home to many species in danger of extinction like the Puerto Rican Parrot. Rain forest trees protect nutrients from eroding and provide important buffers from strong hurricanes. El Yunque is a popular destination for ecotourism, which is tourism that aims to protect the ecosystem. The ecotourism business produces income for Puerto Rico in a way that is less destructive than using the forest as farmland or timber. Humans are adding extra carbon to the atmosphere in the form of carbon dioxide (CO2) by burning fuels like wood, coal, oil, and natural gas. Increased CO2 in the atmosphere is one cause of climate change. The massive trees in El Yunque store large amounts of carbon in their biomass that would otherwise become greenhouse gases in the atmosphere. This important process is called carbon sequestration. The primary concern for tropical forests, including El Yunque, is habitat loss and destruction. Rain forests, and the species that live in them, are being lost rapidly. It is estimated that, worldwide, each hour about 36 km2 of rain forest are cleared. In the same hour, 6 plant or animal species go extinct. The preservation of El Yunque National Forest and the recovery of the Puerto Rican Parrot, which was once on the brink of extinction, are great success stories. Urban areas continue to grow in Puerto Rico and replace natural habitats. Currently only 8 percent of Puerto Rico is protected. It is much easier to protect wildlife on public lands than on private lands. But in Puerto Rico, 90 percent of the island is privately owned. Ecosystem Services • Air quality maintenance: Rain forests play a major role in filtering air pollution, and extracting CO2 from the air through photosynthesis. • Climate Regulation: El Yunque National Forest plays a role in global climate patterns and reduces the negative effects of natural disasters, such as hurricanes and flooding, in the rest of Puerto Rico. • Educational value: The vast biodiversity found within tropical rain forest offers considerable educational and scientific value. • Erosion, flood, and storm control: The tall trees of El Yunque act as a buffer from the strong hurricane winds as storms come ashore. In addition, the root systems keep the precious humus (an important component of soil) from washing away in heavy rainfall. FOSS Populations and Ecosystems Course, Second Edition © The Regents of the University of California Can be duplicated for classroom or workshop use. Version date 12/2015 FOSS Ecoscenarios: El Yunque National Forest General Information Page 3 of 7 • Food: Many popular foods grow in the rain forest. Coffee beans are products of Puerto Rico that grow near the rain forest. • Habitat refuge: Tropical rain forests are home for more than half of the world’s terrestrial species. El Yunque forms the primary habitat for the endangered Puerto Rican Parrot. • Natural Resources: Rain forest trees provide many resources such as timber, paper, and firewood. • Nutrient cycling: Tropical rain forests receive very few nutrients from the outside and must produce most nutrients themselves. When left intact, a rain forest acts as a closed-loop system, recycling the nutrients it has created. • Primary production: Primary production of trees is important for maintaining the carbon cycle. • Soil formation: Trees and plants maintain soil quality by helping produce and hold onto humus (an important component of soil). • Spiritual experience and sense of place: El Yunque offers a deep sense of belonging, cultural heritage, and religious and spiritual significance for many people that visit and live near the forest. • Tourism: El Yunque is a popular destination for ecotourism. Tourism business produces income for Puerto Rico in a way that is less destructive than using the forest as farmland. ABIOTIC DATA Tropical rain forest ecosystems have heavy rainfall and high temperatures year-round. The temperature and length of daylight remain fairly constant throughout the year. There are 11– 13.25 hours of light each day throughout the year. The average air temperature in El Yunque National Forest ranges from 25 degrees Celsius (˚C) to 28°C at lower elevations. The elevations at El Yunque vary from about 100–1000 meters (m). Temperatures are cooler in the mountains, with highs of only 17°C. All these factors provide a year-round growing season for producers. FOSS Populations and Ecosystems Course, Second Edition © The Regents of the University of California Can be duplicated for classroom or workshop use. Version date 12/2015 FOSS Ecoscenarios: El Yunque National Forest General Information Page 4 of 7 It rains daily at El Yunque, with slightly more rain in the winter. Clouds coming from the ocean hit the mountains on the eastern side and drop moisture, while areas to the west of the mountains receive less. Average annual precipitation is 510 centimeters (cm) per year in the forest. San Juan, the capital just west of El Yunque, receives only 150 cm of precipitation per year. Courtesy of National Oceanic and Atmospheric Administration (NOAA) The map shows the distribution of average annual precipitation across Puerto Rico. Notice that the highest levels fall in El Yunque forest. Like most rain forests, El Yunque has clay-like soils that are low in nutrients. Alone, these soils cannot sustain the lush plants found in rain forests. The massive trees and lush plants maintain the soil quality by providing organic material, such as leaves and branches. Decomposers quickly break down this material, creating a nutrient-rich layer of humus that rests on top of the other soil. Tree roots prevent the humus from being washed away by heavy rainfall. When these trees are removed, the humus quickly erodes. Tropical storms and hurricanes frequently pass over Puerto Rico, hitting El Yunque first. Hurricane season in the Caribbean lasts from June to November. High winds and heavy rainfall can damage forests. In 1998, Hurricane Georges was especially destructive. Winds blew over 185 FOSS Populations and Ecosystems Course, Second Edition © The Regents of the University of California Can be duplicated for classroom or workshop use. Version date 12/2015 FOSS Ecoscenarios: El Yunque National Forest General Information Page 5 of 7 km per hour, knocking down many older trees and destroying buildings. The hurricane caused over $5 million in damage. Courtesy of Father Alejandro Sánchez This opening in the canopy is the result of Hurricane George in 1998. In a few years the surrounding palms will close the gap. Sometimes it isn’t the winds that cause the most damage. In 2004, Hurricane Jeanne hit Puerto Rico with relatively mild wind speeds of 130 km per hour. But the storm dumped over 60 cm of rain on parts of eastern Puerto Rico, causing historic floods. Scientists are concerned that climate change may result in stronger hurricanes, followed by more frequent drought events, which could have dramatic effects on El Yunque. BIOTIC DATA Annual primary productivity is the amount of energy provided by the producers to the ecosystem each year. Tropical rain forests like El Yunque have very high annual primary productivity, because the majority of terrestrial producers worldwide are found in rain forests. The tropical climate in El Yunque is ideal for large and lush plant growth. Annual primary productivity is very high in tropical rain forests, like El Yunque, because of the yearlong growing season. FOSS Populations and Ecosystems Course, Second Edition © The Regents of the University of California Can be duplicated for classroom or workshop use. Version date 12/2015 FOSS Ecoscenarios: El Yunque National Forest General Information Page 6 of 7 The rain forest is dense, with a diverse community of producers ranging from tall trees to vines, shrubs, and low-growing plants. Like all tropical forests, El Yunque has four distinct layers: the forest floor, the understory, the canopy, and the emergent layer. The forest floor is covered with leaf litter, detritus, and detritivores, such as termites, cockroaches, and isopods. Leaves that fall on the forest floor may decompose in a few weeks rather than years, because conditions allow for decomposers like bacteria and fungus to flourish. Nutrients are quickly absorbed by nearby plants or washed away by frequent rains. Courtesy of Father Alejandro Sánchez Damp leaf litter decomposes quickly. Courtesy of Father Alejandro Sánchez Centipedes forage for prey on the forest floor. The understory is a mass of vines, dwarf trees like hibiscus, and mushrooms. These producers can survive in low light with very little air circulation. Only about 5 percent of the sunlight filters through to the understory. Plants in the understory rely on insects for pollination. They produce large, sweet-smelling, colorful flowers to attract pollinators. Ants, tree snails, and walkingsticks live in the understory. Predators found in the understory include anoles, geckos, giant crab spiders, tarantulas, and centipedes. The larger trees making up the rain-forest canopy include Sierran palms and Pumpwood trees. The trees have dark leaves to more efficiently absorb sunlight. Rain and moisture that falls on the leaves drop to the forest floor. Epiphytes, like orchids and bromeliads, are plants that grow on other plants without ever touching the ground. Red-tailed hawks and Puerto Rican screech owls make their nests in the tall canopy trees. Endangered species, like the Puerto Rican parrot, live in cavities of larger, older trees. The familiar "koooooooo-keee" call of the common coqui frog can be heard throughout the canopy, understory, and forest floor. The coqui frog is a favorite among Puerto Ricans and is considered an unofficial national mascot. The populations of the Puerto Rican parrot and coqui frog have drastically decreased in Puerto Rico since the 1500s. FOSS Populations and Ecosystems Course, Second Edition © The Regents of the University of California Can be duplicated for classroom or workshop use. Version date 12/2015 FOSS Ecoscenarios: El Yunque National Forest General Information Page 7 of 7 Courtesy of Father Alejandro Sánchez Coqui frogs climb up into trees at night, looking for food and mates. They jump back to the forest floor rather than climb down. The tallest trees make up the emergent layer, which rises above the dense canopy of shorter trees. These trees take advantage of increased air circulation and rely on wind to support pollination and seed dispersal. Thick, waxy leaves reduce the amount of water released through transpiration in the hot, tropical sun. Courtesy of Father Alejandro Sánchez Sierran palms from the forest canopy. Courtesy of Father Alejandro Sánchez Red-tailed hawk and chick in a nest high in the canopy. There are more than 200 species of birds found in Puerto Rico and many more species of insects and invertebrates. Migrating birds depend on resting in places like El Yunque as resting spots along their long journeys. Historical records show 22 kinds of mammals in Puerto Rico. While some still live in Puerto Rico today, such as the Jamaican fruit bat, many of the native species have disappeared. In their place, populations of invasive species, such as black rats and mongooses, now flourish. El Yunque has fewer species than are found in continental tropical rain forests. Islands have a smaller land area for organisms to use, and island populations can become isolated from populations on the mainland. Over time these small populations adapt to their unique environment. When populations of plants or animals are small, there is a greater chance that they will become extinct if too much pressure is placed on the population. FOSS Populations and Ecosystems Course, Second Edition © The Regents of the University of California Can be duplicated for classroom or workshop use. Version date 12/2015