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Announcements Quiz today. Field trip dates Pepperwood- Sun Feb 24 Galbreath- Sat Apr 26 or Sun Apr 27 Darwin Day and Biosystematists Public tours of Essig Museum in afternoon Darwin’s B Day party at 5:00 Biosystematists at 7:30 must submit list today! Evolution of Metamorphosis • • • • Occurred about 300 MYBP in Carboniforous Coincided with evolution of seed plants Two main types Hemimetabolous – Gradual change to adult form – No pupal stage – Includes true bugs • Holometabolous – True larval and pupal stage present – Includes beetles, flies Insects and Plants • Very old associations • Early plant fossils show insect damage • Insects diversified as plants did 1 Diversification in Mesozoic • Diversification of insects that eat plant feeding insects • Appearance of Lepidoptera in Jurassic Appearance of flowering plants in early Cretaceous (130 MYBP) • Diversification of insects associated with flowering plants (Hymenoptera, Lepidoptera) Insect Body Plan (3 regions) 2 ‘Generalized’ insect Insects vary a lot with respect to endoskeleton and appendages Orientation • • • • Dorsal Lateral Ventral Appendages – Proximal – Distal Functions of cuticle • • • • • Forms exoskeleton Prevents water loss Provides support for muscle attachments Lines tracheae and parts of digestive system Varies in toughness – Thick/hard, thin/flexible – Depends on life stage 3 Components of cuticle • Waxes and lipids • Proteins • Chitin – in fungal cell walls and arthropod exoskeletons – unbranched polysaccharide polymer of high MW Structure of cuticle • Epicuticle • Procuticle – exocuticle and endocuticle – made of chitin and protein – Thickest layer • Epidermis – live cells – secretes cuticle 4