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Bell Pettigrew Museum of Natural History Interpretative Panels Text: Dr Iain Matthews Design: Steve Smart & Cavan Convery A University of St Andrews Development Fund Project School of Biology http://biology.st-andrews.ac.uk 15:1 Echinodermata P hy l u m Body Plan: • Bilaterally symmetrical, but appearing 5 rayed • Triploblastic • Spacious coelomic cavity in three parts, and through gut • Deuterostome • Mineral endoskelton with the mesoderm • Complex water vascular system Echinodermata S u b p hy l u m Super class Class The echinoderms are united in having adults with a unique five-rayed (pentamerous) symmetry. Internally, and in their larvae, they show a bilateral ancestry with evidence of a tripartite coelomic body plan. The calcareous endoskeleton is formed from ossicles in the mesoderm, often covered with spines and other protuberances which project through the skin. Gut Endoderm Body Cavity Mesoderm Ectoderm Class: Concentricycloidea Concentricycloidea is the smallest and most recently discovered of the classes of echinoderm. The sea daises are small (<1 cm), disc-shaped echinoderms with podia only around the edges. Only two species, both from deep water (> 1000 m) have so far been found. Another unique echinoderm system, the water vascular system, operates a series of individual tube feet or podia, especially for locomotion. There is no true blood vascular or excretory system, and a very simple radially organised nervous system. Class: Holothuroidea Holothuroidea contains 1150 species of sea-cucumbers. They lack arms, and their elongated, worm-like body lies on its side. The body wall is leathery, with the endoskeleton reduced to A morphologically diverse phylum, the 6750 extant species, spicules. which are all marine and virtually all benthic, are contained Class: Asteroidea in six classes: The 1500 species of asteroid, which includes the starfish and seastars , have flattened bodies with five or more stout arms. Class: Crinoidea Crinoidea contains around 625 extant species of sea lilies and Most species are scavengers or predators with a downward feather stars (e.g. Antedon ). They are usually stalked and pointing mouth and an anus on the upper surface. attached to the substrate with an upward pointing mouth Classification within and a U–shaped gut. Their arms branch repeatedly, forming Echinodermata a filter feeding basket covered with podia and cilia. Class: Echinoidea The 950 species of echinoid, including the sea urchins , and sand dollars , are spherical, possessing large body cavities but lacking arms. Their ossciles are fused into a rigid skeleton, called a test. Echinoids, most of which are benthic detritivores, have a downward pointing mouth, complex jaws (Aristotle’s lantern), and an anus on the upper surface. Class: Ophiuroidea The 2000 species of ophiuroid, which includes the brittlestars and basketstars, are superficially similar to the asteroids. However their five arms are clearly demarcated from the central disc-shaped body. Most species are scavengers having a downward pointing mouth, but lacking an anus. Class: Echinoidea Order: Cidaroida Order: Echinothuroida Order: Diadematoida Order: Pedinoida Order: Salenoida Order: Phymosomatoida Order: Arbacioida Order: Temnopleuroida Order: Echinoida Order: Holectypoida Order: Clypeasteroida Order: Cassiduloida Order: Spantangoida Order: Neolampadoida Order: Holasteroida Class: Holothuroidea Order: Dendrochirotida Order: Dactylochirotida Order: Aspidochirotida Order: Elasipoda Order: Apodida Order: Molpadiida Class: Crinoidea Order: Isocrinida Order: Comatulida Order: Millericrinida Order: Bourgueticrinida Order: Cyrtocrinida Class: Asteroidea Order: Brisingida Order: Forcipulatida Order: Valvatida Order: Notomyotida Order: Paxillosida Order: Velatida Order: Spinulosida Class: Ophiuroidea Order: Oegophiurida Order: Phrynophiurida Order: Ophiurida Class: Concentricycloidea Order: Peripodida See specimen. Brittlestars, sea urchins and sea cucumbers The phylum Echinodermata contains animals with an amazingly wide variety of shapes. It includes the starfish and brittlestars , the sea lilies , urchins and the sea-cucumbers . The echinoderms get their name from the Latin for ‘hedgehog skin’. The crinoids, or sea lilies, now make up only a small part of the phylum. However they are extremely abundant in the fossil record .