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Plant Physiology: How plants work By Adrian Hunsberger University of Florida/IFAS Miami-Dade County Extension What is a Plant? Any of various photosynthetic, eukaryotic, multicellular organisms of the kingdom Plantae. Photosynthetic - the biological process where light energy is converted to chemical energy in the form of a sugar molecule. Eukaryotic - an organism having a nucleus Multicellular - an organism having more than one cell What Makes a Plant? Leaves Flowers Roots Seeds Stems Cellulose Cell Walls Plants - Two Classes Gymnosperms Do not bear fruits, but instead have “naked seeds” in cones. All woody, perennial, usually evergreen. Ex.: Conifers & cycads Angiosperms Have flowers, have seeds which are always in a fruit. Plant Types Annual Short-lived plant. The entire life cycle is completed in one growing season. Biennial Two seasons to complete life cycle Perennial Lives from year to year, either woody or herbaceous. Year 1 Year 2 Plant Growth Plants manufacture their own food (by converting energy from sunlight). All living things require energy, not just for growth and reproduction, but also for the maintenance of life. To produce food the plant requires energy, carbon dioxide, water, and the essential nutrients. How A Plant Grows Photosynthesis - “to put together with light”: light 6CO2 + 6H2O -----------> C6H12O6 + 6O2 Sugars can be transported Starches can be stored Respiration: C6H12O6 + 6O2 ---> 6CO2 + 6H2O + Heat More energy stuff Respiration Breaking the carbohydrates into a form of energy the plant can use Although we think of respiration in animals as breathing, breathing is really the gas exchange that supports respiration. Plant Growth Photosynthesis Produces food Energy is Stored Occurs in Cells with Chloroplasts Oxygen is released CO2 is used Occurs in light Respiration Uses food for Energy Energy is released Occurs in all cells Oxygen is used Water is produced CO2 is produced Occurs in Dark or Light Photosynthesis Occurs only in chloroplasts Chloroplasts contain chlorophyll where photosynthesis occurs Chlorophyll, a plant pigment, appears green Where photosynthesis takes place Photosynthetic Pathway What’s all this used for, anyway? Plant growth and development Cell division Cell elongation Where does growth occur in plants? Meristems Root tip Light • Color – Why don’t plants grow well in green light? - what about violet-blue and red light? Light • Intensity – Shade plants and full sun plants • Duration – Total amount in a day – Photoperiod and flowering Short-day – poinsettia Day-neutral – tomato Long-day - dill Plant Types - Angiosperms Have flowers, have seeds which are always in a fruit Monocots – grasses, palms, bamboo, lilies, orchids, amaryllis Dicots – “broadleaves” How substances move within the plant Phloem – tubes that conduct food and hormones produced by the plant from leaves to entire plant Xylem – tubes that conduct water and minerals from roots to entire plant These tissues are formed by the vascular cambium Woody dicots Herbaceous dicots Monocots Growth - Monocots Cotyledon (seed leaves) = 1 Shoot Elongation Shoots elongate from 1 growing point, pushing material upward. Leaf Growth Occurs from a meristem, a bud, a growing point. Growth - Monocots Stem/root Diameter Growth Stem diameter is determined within the growing point no cambium layer exists. Growth - Dicots Cotyledon (seed leaves) = 2 Shoot Elongation Shoots elongate or grow in height at the tips of the branches. Leaf Growth Occurs from meristem (region making new cells), a bud. Growth - Dicots (Woody) Stem/root Diameter Growth Between the wood and bark is a thin layer of dividing cells, the vascular cambium. Growth - Dicots (Herbaceous) Plant Types Woody- stem tissue is tough and strong (monocots) Stems and roots lengthen each year. (dicots) Stems and roots increase in diameter (thicken) and length each year. Herbaceous Aerial portion of the plant is relatively short-lived and the tissues relatively soft. Growth Factors – plant hormones Very small concentrations of these substances produce major growth changes. There are five groups of plant-growth-regulating compounds: auxin, gibberellin (GA), cytokinin, ethylene, and abscisic acid (ABA). Growth Factors – plant hormones Auxin causes several responses in plants: Bending toward a light source (phototropism) Downward root growth in response to gravity (geotropism) Promotion of apical dominance Flower formation Fruit set and growth Formation of adventitious roots Growth Factors – plant hormones Gibberellic acid (GA) = Gibberellins Stimulates cell division and elongation, breaks dormancy, speeds germination. Cytokinins are found in both plants and animals. Stimulate cell division also are used to delay aging and death (senescence). Growth Factors – plant hormones Ethylene gas Ripening agent Stimulates leaf and fruit abscission. Ethylene is unique in that it is found only in the gaseous form. Plants often increase ethylene production in response to stress Car exhaust contains ethylene! Growth Factors – plant hormones Abscisic acid (ABA) is a general plant-growth inhibitor. It induces dormancy and prevents seeds from germinating; causes abscission of leaves, fruits, and flowers; and causes stomata to close. Growth Factors - Genetic Plant Family Species Cultivar (“variety”) Growth Factors - Environmental Water Air Light Pests Temperature Culture Temperature Heat and cold effect the rate of chemical reactions and therefore, plant growth. Plant Growth Factors Plant Organs - The Root Roots Do not possess nodes, internodes or buds. Function primarily in support, water and mineral uptake from the soil. Grow in both length and diameter as the plant grows (dicots). Commonly grow to 2-3 times the diameter of the shoot (canopy) area; primarily in top 12-18 inches of soil. Plant Organs - The Root Primary Root: generally arises from the embryo as the Radicle (seed root). Tends to be short lived, but may turn into a tap root in some species. Secondary Roots: Branching roots off of the primary root, become the permanent roots. Types of Tree Roots Small absorbing roots Sinker roots Lateral roots Taproot (a temporary structure) Plant Organs - The Root Adventitious Roots - arise from stem structures. Root Hairs - extensions of root cells - these draw all the supplies. Factors Influencing Plant Root Growth Water u Air u Soil temperature u Soil structure u Soil pH u Nutrients u Water Plants are 90% water, most of which is absorbed through the roots Water is a solvent for minerals and translocation Water is used during photosynthesis to manufacture food Water gives cells shape (turgor) Water Extremes With excess water roots will suffocate. Without water most roots dry out and die. Soil Aeration Roots must have oxygen from the air to convert stored food to energy (respiration). Roots need oxygen to absorb essential elements. oxygen carbon dioxide Pores in the soil hold air. Oxygen is most available in the top 18” of soil. Lack of oxygen in the soil may be caused by Planted too deep Saturation Compaction Fill soil Functions of Plant Roots Anchorage Absorption and transport Storage Production of chemicals Anchorage u Roots hold plants upright u Roots keep plants stable as winds blow against them Anchorage Trees need healthy roots for support and stability Tree roots and anchorage fail due to: Excess water Diseased roots Excessive fill or compacted soil Severed roots Absorption and Transport Roots absorb water and nutrients from the soil Roots translocate (move) water and dissolved nutrients to other plant parts Small absorbing roots are covered with fine root hairs Storage Roots store reserves of sugars, starches and other materials Plants use stored reserves for growth and repair Food stored in the roots may be used to produce stems, leaves flowers and new roots. Starch stored in roots stains dark purple with iodine. Knowing where tree roots are located can help us protect them and know where to apply mulch, water, fertilizer and other materials needed to promote root growth • root growth • compacted soil • low soil oxygen • soil erosion • excessive mulch • physical objects Reasons roots occur above ground Some trees (mainly tropicals) have above ground roots for support and absorption. Tree Root Spread Whether a younger or an older tree, roots may extend ... … 2 to 4 times beyond the drip line. Root Spread Limitations (buildings, containers, curbing, median strips, parking islands, tree pits and other obstacles) Plant Organs - The Shoot Consists of stems and leaves. The shoot functions primarily in support, food and water conduction, and food manufacture. Plant Organs - The Stem (Dicots) Nodes Points on a stem where a leaf or leaves are attached. Spaces between nodes are called Internodes. Buds Lateral buds at the base of leaves. Terminal buds at the apex of stems. Adventitious buds on stems. Plant Organs - The Stem The Stem The stem functions primarily in support, food and water conduction, and food manufacture (when green). Sometimes also stores food. Can root at nodes (dicots). The Stem - Monocots Palms, orchids, & grasses are monocots vascular system differs from dicots Plant Organs - The Leaf The Leaf Is a flattened or extended portion of the stem. 2 parts - the blade and the petiole. veins appear as lines or ridges Stomates (stomata) – regulate gas exchange & moisture loss Plant Organs - The Leaf Monocot Leaf Chief veins are parallel or nearly so. Typically grasses, palms Dicot Leaf veins form a net-like pattern. Commonly referred to as “Broadleaves”. Mark Shelby Sarasota County Extension Bruce Ide Palm Beach County Extension Bonnie Lee Appleton Katherine Kilburne Sybil Mays Susan Ruiz-Evans Virginia Polytechnic Institute and State University Hampton Roads Agricultural Research and Extension Center Virginia Beach, Virginia