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Cell types
Sclereids
Sclereids are specialized cells
formed in a variety of shapes.
They have lignified secondary
cell walls and are non-living at
maturity.
Their primary function is for
mechanical
support
and
protection.
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Cell types
Sclereids
Several of the different
sclereid types include:
Brachysclereids
Macrosclereids
Osteosclereids
Astrosclereids
Trichosclereids
Brachysclereids in purple coneflower seed.
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Cell types
Sclereids
Brachysclereids
resemble
parenchyma cells in shape, but
are made almost entirely of
secondary cell wall.
These non-living cells are also
called stone cells because they
form the "grit" in pear fruit.
The inner portion of a sclereid
cell is called the Lumen. The
lumen is an empty space in the
center of a non-living cell
bounded by secondary cell
walls.
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Lumen
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Pits
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Cell types
Sclereids
Macrosclereids are elongated
sclereids usually found in the
outer layer in the seed coat of
legume seeds.
Macrosclereids
These cells are responsible for
restricting water uptake by
hard-seeded legumes.
They are also called Malphigian
cells after the pioneering
Italian 17th century plant
anatomist.
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Seed coat in eastern redbud (Cercis).
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Cell types
Sclereids
If the seed coat is macerated with appropriate chemicals, you can see
the isolated shape of a macrosclereid.
Macrosclereids
Seed coat in eastern redbud (Cercis).
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Cell types
Sclereids
The light line is a continuous line
just beneath the outer surface
of the macrosclereids in some
legume seed coats.
It is actually, an optical
refraction of light due to the
properties of the macrosclereid.
It was once thought to be
responsible for making these
cells impermeable to water, but
this is probably not the case.
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Macrosclereids
Light
line
Seed coat in eastern redbud (Cercis).
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Cell types
Sclereids
Electron micrograph of the seed coat in honeylocust (Gleditsia).
Macrosclereids
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Cell types
Sclereids
Osterosclereids (also called bone
or hour-glass cells) are columnar
cells similar to macrosclereids
except they are enlarged at the
ends making them bone-shaped.
Macerated pea seed coats show
groups of osteosclereids. In side
view, you see the typical "bone shape"
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Cell types
Sclereids
Astrosclereids are branched, pointed, irregular (often star - shaped)
sclereids. Astrosclereids are not a common type of sclereid.
They can be found in specialized tissues like the floating leaves of water lily.
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Cell types
Sclereids
Trichosclereids are uncommon cells.
They can be found in the leaves of
olive and the aerial roots of the
Swiss cheese plant (Monstera sp.).
These long trichosclereids are from
macerated roots of the Swiss
cheese plant.
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