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____________________________ ___________ ________ 0 POPULATIONS A population is a group of organisms of the same species living together in an area. Population density is the number of organisms of a particular species living in an area with a specific size. A population is said to be dense when there are a large numbe organisms living in a relatively small area, and sparse when there are fewer organi r of sms living in the area (or in a larger area). Limits on Populations. Field mice may have litters with six or more pups, and they can reproduce every six weeks, It takes only six weeks for a mouse to become sexually mature. In 6 months, a population of 20 mice could become a population of 5120 mice. Mice have been around for millions of years, so why is it that when you look out the window you see grass and trees, birds and squirrels, and not a vast carpet of millions and millions of mice? The reason is that there are limits on all populations, including those of the prolific mouse. Population Growth Patterns Changes in population size in a community occur when individuals are added to or removed from a population (Table I). If natality (the birth rate) increases while other factors remain constant, the population will increase. The same holds true if immigration increases. The reverse effect occurs if there is an increase in mortality (the death rate) or emigration. In populations in natural ecosystems, all four factors interact with natality and mortality generally having the greatest, effect. The four factors involved in population growth can be expressed mathematically by the formula Population growth = (births + immigration) (deaths + emigration) Biotic Potential One of those limits on population is imposed by the species on itself. Species vary in their capacity to reproduce. iotic potential is the maximum number of offspring that a species could produce, if resources were unlimited. You have seen how quickly field mice reproduce, but many animals have a much lower biotic potential. For example, mature female black bears give birth to one or two cubs after a gestation period of 7.5 months. Generally, bears take at least two years to mature, during which time their mother will not give birth again. Biotic potential isregulared by four important factors, shown in Figure 1. — In mature ecosystems, where resources tend to be constant or available in predictable patterns, populations remain relatively stable over the long term (population growth 0). Increases in nasality are balanced by increases in mortality or increased emigration. This balance is referred to as dynamic equilibrium, or a steady state. Open and Closed Populations Ecologists dassi’ populations as either open or closed, In most natural ecosystems all four factors (nataliry, mortality, immigration, and emigration) are acting on the population of each organism. These populations are said to be open populations. However, immigration and emigration do not happen in laboratory settings and in some game reserves, so populations of organisms in these situations are considered closed populations only narality and mortality affect their population size. — The global population of humans or any other type of organism is also considered a dosed population. People do not emigrate to other planets, and as far as we know none are immigrating either, Changes in the size of a global population result only from natslity and mortality inttt. Factors That Affect Population Size Faster nasality Descriptiso births) mortality (deaths) Immigration emigration the camber of offspring of a species born in sne year the number of individuals of a species that die in one yesr. the number sf isdividualt ste species moving into an existing population. the samber sI individuals of a species moving out of on existing population. iclty tot The ricrher of ng that roach reprodcnve age The female sea turtie sys many eggs butonty stew of her sff$ptisl even reacl the sea, antifewerstil reach rntan.rity,’ C Factsts that determine biotic potential tbdrol ILr. . c,k mate anlycnce per year cfr’the faL Factac tcegth et reprodactive Ill. Deseriptiso: “e age ct seaus ‘aLrt arc tm rr’ber s’ yea’s the ndivitia. ca :eptnce. Afnica eeparrs reach seaus maturty at abOt :1 years stage tt flay rep•’ncuce art.. they are tD. Table I. Factors That Limit Populations cast;. enpsylatIpn Facts$ they cans. e.pnpulaIle to hierakac fctore that 16 decraae favosrable ‘girt favourable remperatarn ‘avourabe chencal envirsrrment too much or too little girt too cold or iso wamr on!avnLrable chemical environment sifficant food sw number or ow effectiveness of insufficient toed high number or hgh effectiveness of pndatnrs predators Ins or weak diseases and parasites abrit to compete for resources many or strnrrg diseases and parasites inability to s’sccassfully rsorpete for resources For example, a fern plant produces more than 50 000 spores in a single year (Figure 2). If all fern spores germinated, fern plants would cover all of North America within two generations of the first plant. This doesn’t happen because of the limiting biotic and abiotic factors. If the weather is wetter than usual, the soil is moist, and many fern spores will germinate, so the fern population will increase. A return to drier weather will not only prevent spores from germinating, but will also kill plants in exposed areas, so the population declines. The presence of many grazing animals will reduce the population of ferrss, and if there are few grazers the population will grow (Figure 2). Fluctuations like these, caused by one factor, can occur in natural ecosystems; however, most populations are affected by more than one factor at a time. Denstty4;depeedeet tacesro • flood • fire • spraying with pesticides • change in climate or temperature • deffructisn of habitat • droaght Densitpdepandeet factors • food shortage • csmpstition for mates, breeding ornas (habitat) • diseasu caassd bya oricrosrganism or parasite • introductiss stan asssic species • increased predation • competitiso fur water and athe, resnarces Populations by answering the following Read the accompanying pages. Prepare your own notes e part of the question in the (includ le possib er S questions in FULL SENTENCE wherev answer to make complete, detailed notes): 1. a) Define population, population density. two? Which do you think is b) What seems to be the main difference between these more useful to know, and why? 2. a) Define the following terms: 3) immigration 1) natality 4) emigration 2) mortality . b) Give the mathematical formula for population growth tems seem to be like. What 3. Describe what populations In mature (established) ecosys are the reasons for this? tions. 4. a) Distinguish clearly between open and closed popula each? in size tion popula the ine determ to tend factors b) Which 5. a) What is the blotic potential of a population? al of a population. b) Describe the 4 factors that determine the biotic potenti biotic potential of a population? the it affect does how and factor” “limiting is a What 6. a) s ablotic and biotic factors can b) Recreate Table 1 in your notes to show how variou limit a population. 7. a) Define carrying capacity of a population. tion? b) What determines the carrying capacity of a popula what happens when a explain tion, popula mouse field the of le examp c) Using the tem. ecosys the ty of capaci g carryin population exceeds the factors and density-dependent 8. a) Distinguish ciearly between density-independent le to illustrate each. examp e an Includ tions. popula affect can that factors of these types of factors. each of les examp list to notes In your b) Recreate Table 2 ITh Abintic and bistir factors limit tha nsmber of fems in an ecosystem. Carrying Capacity Populations fluctuate regularly due to an interaction of the many biotic and abiotic limiting factors. However, communities do tend toward stability. Stability is achieved when an ecosystem is in equilibrium, when none of the populations exceeds the carrying capacity of the ecosystem. The carrying capacity is the maximum number of individuals of a species that can be supported indefinitely by an ecosystem. The carrying capacity for any species is determined by the availability of resources, such as food and water. A population can exceed the carrying capacity of the ecosystem, but not for long. Consider the field mouse again. Imagine that the population of predators is lower than usual. Suddenly, the mouse population can grow. However, the extra mice will eat all the available food. Hungry rodents aoon become sickly making them easy prey for the hawks, owls, and foxes that are present. The mouse population will decline again, to or below the carrying capacity. Ecosystems soon re-establish equilibrium. — c. a;t ering The number of organisms in an ecosystem is important when consid the effects of some abiotic and biotic factors. A population is said to be dense when there is a large number of organisms in a small area. Density-independent factors affect members of a population regardless of population density. Fire and flood are two naturally occurring events that are density-independent. They will affect a population regardless of its size. When the density of a population increases, other factors may limit further growth or reduce population numbers. Density-dependent factors affect a population because of the density of the population. Food supply, water quality sunlight, disease, and territory are density-dependent factors. For example, when a tree in a dense forest becomes infected with a fungsl blight, the infection will spread more quickly than it would in a forest where trees are separated by larger distances. Similarly, individuals in more densely populated areas are more prone to starvation, as food is in lower supply. Competition for food may leave animals weak and more susceptible to predation. The density-dependent factors in Table 2 will cause higher ynortality rates, lowering the population density When the population density is reduced, the effects of the density-dependent factors are also reduced. ng The environment provides factors that prevent populations from attaini their biotic potential. Any resource that is in short supply is a limiting factor on a population. Food, water, territory, and the presence of pollutants and other toxic chemicals are all limiting factors, as shown in Siatto Factors that Cause Chances in Populations Density Dependent and Independent Factors Limiting Factors Ab)ntlc G 0 0 J)