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Growth Movements - tropisms Plants may not be able to get up and go, but they do move. Plants can change the angle of their leaves so that the leaves track the movement of the sun in the sky. Plants can bend their stems toward the light, too, and their roots grow downward in response to gravity. Some plants even have quick movements, like those of the Venus flytrap, which enable them to respond to the presence of predators and prey. Growth Movements - tropisms One type of plant movement – a tropism – is a directional movement that results from growth in a particular direction. These growth movements occur in response to elongation of selected cells. Plants exhibit tropic growth in response to many signals, including light, gravity, chemicals, and temperature. Growth Movements - tropisms Phototropism: If light shines on a plant from one side, the plant shoot will exhibit positive phototropism, and bend toward the light. If you’ve ever grown plants near a window, you’ve probably seen evidence of this particular tropism. Molecules called photoreceptors in the tips of the plant shoots absorb blue light. Growth Movements - tropisms Signal transduction occurs, leading to a greater production of auxin on the dark side of the shoot. Auxin promotes cell elongation, so the cells on the dark side elongate faster than the cells on the light side and the shoot bends toward the light. Growth Movements – tropisms Growth Movements - tropisms Geotropism (gravitropism): Plant roots grow toward the pull of gravity, so they show positive geotropism. Plant shoots, on the other hand grow away from gravity, showing negative geotropism. You can test this by laying a plant on its side. After awhile, the shoots will turn and grow upward, while the roots will turn and grow downward. Growth Movements - tropisms Growth Movements - tropisms A great deal of evidence supports explanation for geotropism outlined by the starch-statolith hypothesis. According to this hypothesis, plants sense the pull of gravity on amyloplasts located in the cells of the root cap and close to the vascular bundles in shoots. The amyloplasts get pulled by gravity so that they collect on one side of the cell that contains them. Growth Movements - tropisms Auxin is produced in response to the pressure of the amyloplasts, and transported to the side of the root or shoot where the amyloplasts have accumulated. In shoots, auxin stimulates cell elongation, so the side of the shoot where the amyloplasts have accumulated grows faster, and the shoot curves up. Growth Movements - tropisms In roots, high concentrations of auxin actually inhibit cell elongation, so the side of the root with the amyloplasts grows more slowly than the other side. The root curves downward. Growth Movements - tropisms Thigmotropism: This tropism is in response to the plant touching a thing. One of the most familiar examples of thigmotropism is when a climbing plant, such as a pea or a vine, curls a tendril around a support. Because these plants curve toward the solid surface, they exhibit positive thigmotropism. Growth Movements - tropisms Roots, on the other hand, will curve away from solid objects like the rocks they encounter in the soil, so they exhibit negative thigmotropism. The mechanisms for thigmotropism are still being investigated, but some things are known. Some extremely rapid curvature responses result from changes in turgor pressure. Growth Movements - tropisms Prolonged curvature results from differential growth, where one side of the shoot or root grows more rapidly than the other. The touch signal is received by epidermal cells, and then signal transduction results in differential growth of the shoot or root. Unlike phototropism and gravitropism, thigmotropism does not seem to be controlled by auxin. Growth Movements - tropisms Turgor Movements Turgor movements occur in response to changes in turgor pressure in certain cells. Whereas growth movements take some time because the plant actually has to grow, turgor movements are very rapid. For example, some tendrils exhibit rapid coiling around a solid support, positive thigmotropism, due to turgor movements, followed by continued curved growth due to growth movements. Turgor Movements During turgor movements, cells rapidly lose their turgor pressure and collapse, causing the plant to fold around those cells. Here’s what scientists think is happening to cause these responses in tendrils: 1. Touch signals sense the pressure of a solid object. 2. Signal transduction occurs, causing cells near the touch stimulus to export potassium ions outside the cells, thus making the environment hypertonic to the plant cells. Turgor Movements 3. Water leaves the plant cells, following the potassium ions, and the cells collapse, causing the plant to fold. In addition to curving tendrils, turgor movements also occur in Mimosa pudica, which is sometimes called the sensitive plant. If you run your fingers along the leaves of this plant, they fold up immediately. Turgor Movements At the base of each leaflet is a swollen structure called a pulvinus. When you touch the leaves of the sensitive plant, the signal is relayed to the pulvinus. Cells on one side of the pulvinus lose their turgor pressure, causing the leaves to fold together.