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The Cenozoic Era Geology 103 Increasingly Familiar • Following the Mesozoic extinctions, – marine invertebrates diversified giving rise to the present-day familiar marine fauna • Overall, we can think of the Cenozoic Era – as a time during which Earth's flora and fauna became increasingly familiar • Cenozoic rocks are more easily accessible – at or near the surface, so we know more about Earth and life history for this time than for any previous eras. Good Fossil Records • Cenozoic rocks are especially widespread in western North America – although they are also found along the Gulf and Atlantic coasts • As a result, we have a particularly good fossil record for many organisms – Several of national parks and monuments in the West feature displays of fossil mammals, including: • • • • • Agate Fossil Beds National Monument in Nebraska, Badlands National Park in South Dakota, John Day Fossil Beds National Monument in Oregon Hagerman Fossil Beds National Monument in Idaho La Brea Tar Pits in Los Angeles John Day Fossil Restoration • Restoration of Clarno Formation fossils – from Eocene epoch rocks in John Day Fossil Beds National Monument, Oregon – The climate at this time was subtropical and the lush forests of the region were occupied by early rhinoceroses tapirs Titanotheres standing 2.5m high at the shoulder ancient horses carnivores La Brea Tar Pits • Restoration of a Pleistocene mammoth trapped in the sticky tar at a present-day oil seep at least 230 kinds of animals were found trapped in the sticky residue, including dire wolves and sabre-toothed cats Diatoms • Diatoms were particularly abundant – during the Miocene, probably because of increased volcanism during this time • Volcanic ash provided increased dissolved silica in seawater which diatoms used to construct their skeletons • Massive Miocene diatomite is present in several western states Diatomite • Outcrop of diatomite from the Miocene – Monterey Formation, Newport Lagoon, California Foraminifera • The foraminifera were a major component of the Cenozoic marine invertebrate community • Although dominated by relatively small forms (microscopic), it included some exceptionally large forms (naked-eye visible) that lived in the warm waters of the Cenozoic Tethys Sea • Shells of these larger forms accumulated to form thick limestones, some of which ancient Egyptians used to construct the Sphinx and the Pyramids of Giza Cenozoic Foraminifera • A planktonic form – Globigerinoides fistulosus – Pleistocene, South Pacific Ocean Corals—Reef Builders Again • Scleractinian corals were perhaps the main beneficiary of the Mesozoic extinctions • Having relinquished their reef-building role to rudists during the mid-Cretaceous, corals again became the dominant reef builders • They formed extensive reefs in the warm waters of the Cenozoic oceans and were especially prolific in the Caribbean and IndoPacific regions Evolution of Mammals • Mammals evolved during the Late Triassic, – and some Mesozoic mammals retained characteristics of their ancestors, the cynodonts. • By Cenozoic time, mammals had clearly differentiated from their ancestors. Class Mammalia • All warm-blooded vertebrates with hair and mammary glands are members of the class Mammalia, – which includes two fundamentally different kinds of mammals, the prototheria and the theria (or eutheria) • The prototheria include some extinct animals – but the only living ones are the monotremes • order Monotremata, or egg-laying mammals – the platypus and spiny anteater of the Australian region Therian Mammals • Therians, in contrast, include all mammals – that give birth to live young – such as marsupial mammals • order Marsupialia, • commonly called “pouched mammals” – and the placental mammals • with about 18 living orders Uintatheres • Scene from the Eocene – showing the rhinocerossized mammals – known as Uintatherium • They had three pairs – of bony protuberances on the skull – and saberlike upper canine teeth Megaloceros • Giant deer Megaloceros giganteus – commonly called Irish Elk – lived in Europe and Asia during the Pleistocene – Large males had antler spreads of nearly 4 m Other Biological Events • Angiosperms (flowering plants) continued to dominate land plant communities and now constitute more than 90% of all land plants • Birds evolved during the Jurassic, perhaps earlier – the families now common appeared during the Paleogene and Neogene, reached their maximum diversity during the Pleistocene Epoch, and have declined slightly since then What periods are in the Cenozoic? The basic problem is that there aren’t many “big” mass extinctions in this era! A couple of schemes for subdividing the Cenozoic are shown but the extinctions are not as major as in the Mesozoic This lack of clearly-defined boundaries leads to nomenclature confusion; the Encyclopedia Britannica states: “The term Paleogene was devised in Europe to emphasize the similarity of marine fossils found in rocks of the first three Cenozoic epochs, as opposed to the later fossils of the Neogene Period (23 million to 2.6 million years ago). In North America, the Cenozoic has traditionally been divided only into the Tertiary Period (66 million to 2.6 million years ago) and the Quaternary Period; however, the notion that the Tertiary should be replaced by the designations Paleogene and Neogene is becoming more widespread.” Anyhow, the Zuni transgression ends and the Tejas transgression begins Antarctica separates from Australia, heads to the South Pole (Eocene) Paleocene-Eocene thermal maximum • Around 55 my ago, temperatures worldwide increased to levels similar to the early Mesozoic • Cause is uncertain, but may have involved increased atmospheric methane due to warming of ocean floor releasing methane from methane clathrates • Caused many reptilian and mammalian orders to go extinct PETM deep ocean temperatures were high So life survived global warming without too much fuss; why is the current version different? Leaf Structure • Leaf structure is a good climatic indicator • For instance, leaves with entire or smooth margins, – many with pointed drip-tips, – are found mostly in areas with abundant rainfall – and high annual temperatures • Smaller leaves with incised margins – are more typical of cooler, drier areas Plant leaves as Climatic Indicators • Fossil floras with mostlysmooth-margined leaves with drip-tips indicate the climate was wet and warm • while small leaves with incised margins indicate a cool, dry climate Paleocene Flora • Paleocene rocks – in North America's western interior have fossil ferns and palms, both indicating a warm subtropical climate • In a recently discovered Paleocene flora – in Colorado with about 100 species of trees, nearly 70 percent of the leaves had smooth margins and many had drip tips • This range of diversity is much like that found in today's rain forests Decrease in Precipitation • A general decrease in precipitation – over the last 25 million years took place in the midcontinent region of North America • As the climate became drier, – the vast forests of the Oligocene gave way first to savanna conditions • grasslands with scattered trees – and finally to steppe environments (short-grass prairie of the desert margin) Climate story of the Cenozoic is cooling Grasses: a new type of photosynthesis C4 plants evolved in the Oligocene as a response to lowering CO2 levels, drought conditions and other environmental stresses. Arose in many families of plants simultaneously; a good example of convergent evolution Whales: another study of convergent evolution • Eocene hippo-like terrestrial mammals to Oligocene marine mammals • Evolution of whales stumped Darwin • Molecular biology and discovery of transitional forms helped explain history Whale Cladogram • Cladogram showing the relationships – among some fossil and living whales, and their land-dwelling ancestors • Pakicetus had well-developed hind limbs, – but only vestiges remain in Protocetus and Basilosaurus Cope’s rule • Edwin Drinker Cope (1870) proposed this rule to describe the changes in size of organisms as they evolved • As time progresses, organisms of the same clade increase in size • Horses over the Cenozoic are a good example Cooler Conditions—Larger Sizes • Of course, many smaller mammal species – also existed, but one obvious trend among Pleistocene mammals was large body size • Perhaps this was an adaptation to the cooler conditions that prevailed during that time • Large animals have less surface area compared to their volume – thus retain heat more effectively than do smaller animals Himalayan orogeny occurs in Pliocene and continues today • India detaches from Gondwana and heads north • Collides with Asia about 10 my at a (tectonically) rapid rate • Raises Himalayas, Tibetan Plateau, Tien Shan • Causes northern subtropical jet stream to split around the uplift, and repositions the high and low pressure areas over the Pacific Cenozoic climate trend is drier 5.96 my – Strait of Gibraltar closes 5.96 – 5.33 my – Messinian salinity crisis 5.33 my – Zanclean flood event Closing of Panama isthmus • Subduction-related volcanism closes isthmus about 3 my • Allowed ice sheets in Northern hemisphere to form • More salty water in Atlantic • Water sinks because it cools as it heads north • Releases heat as it sinks, so polar oceans never get warmer water Ice ages start up about 2 my ago • Why then? • Several factors need to be in place: polar continent, north/south-oriented oceans, split in the sub-tropical jet stream • At that point, subtle factors can determine whether the glaciers build up or not Milankovitch cycles • Milutin Milankovitch (University of Belgrade, Serbia) proposes that “orbital parameters” that vary in a cyclic fashion over tens of thousands of years will determine that amount of solar insolation that reaches the Earth’s surface Evidence of Pleistocene glaciations End moraines are mounded till deposits that represent the furthest advance of a glacier Pleistocene Extinctions • During the Pleistocene, the continental interior of North America was teeming with horses, rhinoceroses, camels,mammoths, mastodons, bison, giant ground sloths,glyptodonts, sabertooth cats, dire wolves, rodents, and rabbits • Beginning about 14,000 years ago, many of these animals become extinct, especially the larger ones. • Why? Scientists are currently debating two competing hypotheses: Rapid climate change vs. Prehistoric overkill Pleistocene Extinctions • These Pleistocene extinctions were rather modest compared to those of the Paleozoic and Mesozoic eras, but they did have a profound effect on large land-dwelling mammals. • Particularly hard hit were Australia and the Americas • In Australia, 15 of the continent's 16 genera of large mammals died out • North America lost 33 of 45 large-mammal genera, and South America lost 46 of 58 such genera • In contrast, Europe lost only 7 of 23 such genera, and in sub-Saharan Africa only 2 of 44 died out Climate and Vegetation Changes • Rapid changes in climate and vegetation occurred over much of Earth's surface during the Late Pleistocene, as glaciers began retreating • The North American and northern Eurasian open-steppe tundra were replaced by conifer and broadleaf forests as warmer and wetter conditions prevailed Climate and Vegetation Changes • The Arctic region flora changed from a productive herbaceous one that supported a variety of large mammals, to a relatively barren water-logged tundra that supported a far sparser fauna • The southwestern U.S. region also changed from a moist area with numerous lakes, where saber-toothed cats, giant ground sloths, and mammoths roamed, to a semiarid environment unable to support a diverse large mammalian fauna Why Didn't Large Mammals Migrate? • For example, reindeer and the Arctic fox lived in southern France during the last glaciation and migrated to the Arctic when the climate became warmer Lack of correlation between extinctions and the earlier glacial advances and retreats • Previous changes in climate were not marked by episodes of mass extinctions Arrival of Humans • Proponents of the prehistoric overkill hypothesis, argue that the mass extinctions in North and South America and Australia coincided closely with the arrival of humans • Perhaps hunters had a tremendous impact on the faunas of North and South America about 11,000 years ago because the animals had no previous experience with humans • The same thing happened much earlier in Australiasoon after people arrived about 40,000 years ago Scattered Communities • No large-scale extinctions occurred in Africa and most of Europe, because animals in those regions had long been familiar with humans • One problem with the prehistoric overkill hypothesis is that archaeological evidence indicates the early human inhabitants of North and South America, as well as Australia, probably lived in small, scattered communities, gathering food and hunting, concentrate on smaller, abundant, and less dangerous animals Though X-ray evidence from Manis, WA, shows otherwise… Clovis point spear tip imbedded in a mammoth femur Indirect Effects of Past Glaciation • Large pluvial lakes (formed in a period of abundant rainfall) existed in closed basins in Utah, Nevada and eastern California – Great Salt Lake is a remnant of the much larger pluvial Lake Bonneville – Huge floods emanated as icedammed lakes (e.g., Lake Missoula) drained catastrophically • Sea level was significantly lowered by large amounts of water locked up into ice sheets, allowing stream channels and glaciers to erode valleys below presentday sea level Giant gravel ripples formed during draining of Lake Missoula The Missoula Floods – 13,000 years ago DRY FALLS by John Knapp http://www.bmi.net/knapp/whitman.html May have occurred up to 90 times as ice dam rebuilt and broke OKANOGAN LOBE Dry Falls is 3.5 miles wide with a drop of over 400 ft. Summary of the flood