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Mollusks and Annelids Chapter 45 Copyright © McGraw-Hill Companies Permission required for reproduction or display Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Coelomates • Coelomate Body cavity lined with mesoderm cells Allows the development of complex tissues and organs. Allows wider array of body architectures and increased body size. Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Coelomates • Mollusks (Mollusca) Extremely diverse. Characterized by a coelom. - Great economic significance Pearls Mother of Pearl - Economic / Environmental Costs Zebra Mussel Invasion Intermediate hosts for parasites. Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Body Plan of the Mollusks • • • • Distinct bilateral symmetry. Digestive, excretory, and reproductive organs all concentrated in a visceral mass and a muscular foot. May have differentiated head. Folds constituting a mantle. Gills - Increased surface area for gas exchange. Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Body Plan of the Mollusks • • • • Shells serve primarily for protection. Radula - Rasping tongue-like organ used for feeding. Circulatory system (except cephalopods) consists of a heart and an open circulatory system. Nitrogenous wasted removed by nephridia. Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Body Plan of the Mollusks • Reproduction in Mollusks Most have distinct male and female individuals. Most engage in external fertilization. Many have free-swimming larvae (trochophores) which closely resemble larval stage of many marine annelids. - Veliger stage follows trochophore stage. Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Classes of Mollusks • • Polyplacophora: The Chitons Oval bodies with eight overlapping calcareous plates. Gastropoda: Snails and Slugs Heads of most have pair of tentacles with eyes at the ends. Undergo torsion during embryological development. Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Classes of Mollusks • Bivalvia: The Bivalves Clams, scallops, mussels and oysters. Have two lateral shells hinged together dorsally. Mantle secretes shell and ligaments. Most are sessile filter-feeders. Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Classes of Mollusks • Cephalopoda: Octopuses, Squids, Nautilus Most intelligent of the invertebrates. Active marine predators. Foot evolved into a series of tentacles equipped with structures to capture prey. Highly developed nervous systems. Closed circulatory system Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Segmented Animals • • Building of body from series of similar segments. Small change in existing segment can produce new kind of segment with different function. Annelids Three characteristics: - Repeated Segments - Specialized Segments - Connections Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Segmented Animals • Body Plan of the Annelid Tube within a tube. Internal digestive tract within the coelom. - Specialized for different functions. Hydrostatic skeleton for locomotion. Each segment typically possesses setae, that help anchor during locomotion. Most have closed circulatory system. Nephridia collect and transport wastes. Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Classes of Annelids • • Polychaeta: The Polychaetes Well developed head with specialized sense organs. Parapodia on most segments. Usually lack permanent gonads. Oligochaeta: The Earthworms Hermaphroditic Copyright © McGraw-Hill Companies Permission required for reproduction or display Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Classes of Annelids • Hirudinea: The Leeches Occur mostly in fresh water. Hermaphroditic Develop clitellum during breeding season. Unable to self-fertilize. Secrete anticoagulant into wounds. Copyright © McGraw-Hill Companies Permission required for reproduction or display Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Lophophorates • Phylum Phoronida: The Phoronids Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Lophophorates • Phylum Ectoprocta: The Bryozoans Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Lophophorates • Phylum Brachiopoda: The Brachiopods Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Review • • • • • • Coelomates Body Plan of the Mollusks Classes of Mollusks Segmented Animals Classes of Annelids Lophophorates Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies Copyright © McGraw-Hill Companies Permission required for reproduction or display Raven - Johnson - Biology: 6th Ed. - All Rights Reserved - McGraw Hill Companies