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Community Ecology Community Ecology Community interactions: Community Ecology Community interactions: –Competition –Predation –Herbivory –Parasitism –Disease –Mutualism –Commensalism –Coevolution Community Ecology Community interactions: –Competition –Predation –Herbivory –Parasitism –Disease –Mutualism –Commensalism –Coevolution -/+/+/+/+/+/+ +/0 Community Ecology Community structure: Species diversity Community Ecology Community structure: Species diversity How many species are in a community? Community Ecology Community structure: Trophic structure Who eats whom? Community Ecology Community structure: Primary producers Community Ecology Community structure: Primary consumers Primary producers Community Ecology Community structure: Secondary consumers Primary consumers Primary producers Community Ecology Community structure: Tertiary consumers Secondary consumers Primary consumers Primary producers Community Ecology Quaternary consumers Community structure: Tertiary consumers Secondary consumers Primary consumers Primary producers Community Ecology Community structure: Energetic hypothesis: Length of a food chain is limited by inefficiency of energy transfer Community Ecology Community structure: Energetic hypothesis: Length of a food chain is limited by inefficiency of energy transfer Dynamic stability hypothesis: Long food chains are less stable than short food chains. Community Ecology Community structure: Energetic hypothesis: Length of a food chain is limited by inefficiency of energy transfer Dynamic stability hypothesis: Long food chains are less stable than short food chains. Community Ecology Community structure: Species with a large impact Community Ecology Community structure: Species with a large impact •Dominant •Keystone •Foundation Community Ecology Community structure: Species with a large impact •Dominant – most abundant •Keystone – disproportionate impact •Foundation – ecosystem engineers Community Ecology Community structure: Bottom-up vs. top-down controls Community Ecology Community structure: Bottom-up vs. top-down controls Predators depend on prey vs. Predators control prey Community Ecology Community structure: Disturbance Community Ecology Community structure: Disturbance May be the norm Community Ecology Community structure: Disturbance Moderate disturbance may foster increased diversity Community Ecology Community structure: Disturbance Human disturbance can be too high or sometimes too low. Community Ecology Community structure: Ecological succession Community Ecology Community structure: Ecological succession •Primary – starting with bare rock •Secondary – starting with soil Community Ecology Community structure: autotrophic prokaryotes Community Ecology Community structure: autotrophic prokaryotes lichens and mosses Community Ecology Community structure: autotrophic prokaryotes lichens and mosses grasses and shrubs Community Ecology Community structure: autotrophic prokaryotes lichens and mosses grasses and shrubs trees Community Ecology Community structure: Pioneer species Climax species Community Ecology Community structure: Pioneer species •Facilitate later species Climax species •Inhibit later species Community Ecology Community structure: births - deaths r = ----------------------N Community Ecology Community structure: births - deaths r = ----------------------N ΔN ------- = rN Δt Community Ecology Community structure: births - deaths r = ----------------------N ΔN K-N ------- = rN (-----------) Δt K Community Ecology Community structure: r-strategies K-strategies Community Ecology Community structure: r-strategies •Rapid growth without competition K-strategies •Maxium population with crowding