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Teufel colors We have developed 16 colors that are isoluminant, equally detectable and perceptually equidistant. They cover the hues available on a standard color monitor and provide a convenient representation for illustrating a difference between receptoral and postreceptoral mechanisms. The position of these colors in the space spanned by the excitations of S-, M- and L-cones (cone space) has been determined using psychophysical methods such as flicker fusion photometry and detection experiments. In conventional cone space (MacLeod & Boynton 1979) these colors form an ellipse situated on a plane of equal luminance. Such ellipses were described first by MacAdam (1942) plotting his results in CIE space. Luminance is often defined as the sum of L- and M-cone signals. Using flicker fusion photometry we first determined the plane of equal brightness for 5 color normal observers. Due to a small contribution of S-cones to the perceived brightness the plane of equal brightness is tilted with respect to the sum of L- and M-cone signals (see Figure a). Generally, scaling of the three base vectors of cone space (the cone fundamentals) is set arbitrarily to 1 (Kaiser & Boynton 1996). Rescaling cone fundamentals with s = .06 m = .37 l = .63 yields the simplest representation of colors using the polar coordinates r (radius) and (azimuth angle) in cone space or cone difference space. Equally detectable colors lie on a circle with radius r being equal to the detection threshold in a plane of equal luminance in cone difference space (see figure a). Increase of azimuth angle corresponds to going counterclockwise around the circle. The 16 colors shown at the end of this page are generated by advancing in steps of equal increments of around a circle with a radius of five times the detection threshold (as measured in five observers). Following the thin arrows in the figure - going from a to b illustrates how the cone excitations of two of the colors situated at azimuth angles 180° and 270° are transformed into S-cone contrasts (cS), M-cone contrasts (cM) and L-cone contrasts (cL) at corresponding azimuth angles . To test the validity of this representation the results of many detection experiments of five subjects are plotted (see figure at the right hand side above). Observers had to indicate the occurrence of small rectangular chromatic and isoluminant stimuli to the left or right of the fixation point. Thresholds were computed at 75% of psychometric functions. Each point (cS, cM, cL) is the result of 300 stimulus presentations. These data are well approximated by the cone contrast functions shown in b. The best-fitting sinusoidal functions in cone contrast space that approximate the detection circle are cS = 0.1006 sin ( - 289.1) cM = 0.0098 sin () cL = - 0.0040 sin ( - 48.8). These functions are shown in the figure (panels a – c) below. They describe the contributions of the three receptoral mechanisms to the detection of the respective colors indicated by stars. Using cS, cM and cL the contributions to the detection of the 16 colors can be expressed in terms of the postreceptoral chromatic mechanisms blue- yellow { cS- (cM+ cL)} and red-green {cM - cL}. These are are shown in panels d and e in the figure below. Since the colors are isoluminant, the receptoral mechanisms S-cone contrast plotted in a. and the post receptoral mechanism blue-yellow plotted in d. are indentical for the 16 colors considered here. We use these colors to study percepts in human observers, for example the shift in color of a gray test field surrounded by either of the 16 colors displayed on a color monitor. The three monitor guns (R,G,B) are calibrated with respect to their spectral radiances. Psychophysical thresholds can then be transformed into values of c S, c Mand c Land plotted as a function of the azimuthal angle Φ. From these the contributions of the postreceptoral mechanism red-green {c M-c L} can be computed. We can predict the outcome of experiments for a range of hypotheses related to the chromatic mechanisms. As mentioned above use of these colors in psychophysical experiments requires a calibrated monitor and an otherwise well defined setting. Monitors used for browsing the internet generally do not satisfy these conditions. The picture below qualitatively mirrors their detectability and hue range. The CIE coordinates of these colors are published in Teufel & Wehrhahn (2000). In this paper also their derivation from flicker fusion photometry and detection experiments as well as geometrical considerations of a color space similar to that of the MacLeod & Boynton (1979) cone space can be found. Literature: Kaiser PK Boynton RM (1996) Human Color Vision. Optical Society of America, Washington D.C. MacAdam DL (1942) Visual sensitivities to color differences in daylight. J Opt Soc America 32: 247-274 MacLeod DIA Boynton RM (1979) Chromaticity diagram showing cone excitation by stimuli of equal luminance. J Opt Soc America 69: 1183-1186 Teufel H Wehrhahn C (2000) Evidence for the contribution of Scones to the detection of flicker brightness and red-green. J Opt Soc America A 17: 994-1006