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Phylum Mollusca November 3-4, 2014 Chiton Conch Clams, Oysters Nudibranch Octopus Squid General Characteristics Soft-bodied invertebrates Unsegmented body, typically bilateral symmetry Internal or external calcium carbonate shell (usually) Second largest animal phylum ~ 200,000 species Show Cephalization Sensory organs towards one end of body “head” formation Habitat/Lifestyle Sessile-mobile Benthic-pelagic Predators-Herbivores-deposit feeders-filter feeders Typical Body Plan of Mollusks Three main parts: Foot Mantle Shell Foot: On ventral side muscular; used for crawling, burrowing, or capturing prey Can be flat (snails) spade-shaped (clams) or modified into tentacles (squid and octopus) Abalone Body Plan of Mollusks Body covered with protective mantle: Thin layer of tissue that covers most of the mollusk’s body May or may not produce a shell Shell: made by glands in mantle that secrete calcium carbonate Feeding with Radula Diverse: herbivores, carnivores, filter feeders, detritivores. Some use Radula for feeding Radula: flexible, tongue-shaped structure covered with tiny teeth Herbivores use radula to scrape or cut algae; carnivores use radula to drill through shells or flesh. Radula Filter Feeding Clams, oysters, and scallops filter feed using feathery gills Mucus produced by gills traps plankton as water comes in through siphons Mollusc Reproduction & Development Sexual reproduction, external In tentacled species, fertilization is internal and female lays fertilized eggs Most have separate sexes that cross-fertilize eggs Some hermaphrodites-both sexes Larval stages Trochophore larva Free swimming, ciliated, shell formation begins Veliger larva Free swimming Veliger Zygote Trochophore Juvenile Adult Spat Respiration Aquatic species use gills inside mantle to exchange gases Incurrent and excurrent siphon brings water in Siphon: Tube-like structure through which water enters and leaves Circulation Can be open or closed circulatory system Open system: blood is pumped by simple heart into sinuses (large saclike space) then to gills, then back to heart; slower system; found in snails and clams Closed system: blood is pumped by heart through blood vessels; faster system; found in faster-moving mollusks 6 Classes in Phylum Mollusca 1)Monoplacophora-simple hat shaped shells 2)Polyplacophora----chitons, 8 overlapping shells 3)Scaphopoda-tusk shells ------------------------------------------------------------------------- 4) Gastropoda-snails, slugs, nudibranchs, and conchs, limpets, abalone 4) Bilvalvia - clams, oysters, scallops, and mussels 5) Cephalopoda squid, octopus, and nautilus Class Gastropoda “Stomach footed” molluscs Have a single shell (snails) or no shell (slugs) Class Gastropoda NUDIBRANCHS Marine slug Lacks shell Class Gastropoda NUDIBRANCHS Feed on sponges Cnidaria, algae Exposed feathery gills Can be toxic Retain undigested nematocysts from Cnidaria=Kleptoplasty Class Bivalvia, Bi-valve = two shells two-part, hinged shell Ex: clams, oysters, scallops mussels Most sessile, benthic Most are filter feeders No head or radula Burrow Sand, wood, rocks A few can use muscular foot that extends from shell for movement Scallops clap valves together to move Class Bivalvia Shell secreted by mantle Oldest, raised part of shell called umbo Have an incurrent & excurrent siphon that circulate water over the gills to remove food & oxygen Have heart & open circulatory system Class Cephalopoda Octopus, squid, cuttlefish, & chambered nautilus Move by using mantle cavity as jet propulsion Sucks water into mantle cavity, pushes water out of muscular siphon that can change size (speed) and direction. Most intelligent mollusk Well developed head –centralized brain Large eyes, visual predators Lack outer shell, very flexible Active, free swimming predators Foot divided into tentacles with suckers Class Cephalopoda Chromatophores: can change color, used for camoflauge and social signaling. 1st group with social signaling-reguires more complex brain to interpret signals Use their radula & beak to feed Closed circulatory system Highly developed nervous system with vertebrate-like eyes Sexual reproduction Separate sexes with internal fertilization Class Cephalopoda SQUID Largest invertebrate is the Giant Squid Large, complex brain 8 tentacles 2 arms- longer pair to catch prey Use jet propulsion Chromatophores Have internal shell called pen Class Cephalopoda Can squirt an inky substance into water to temporarily blind predators Class Cephalopoda OCTOPUS Eight tentacles Similar to squid Sit & wait predators = ambush Female dies after tending to eggs. Beak like jaws, radula Importance of Mollusks Filter water, healthy waters Keep algae growth done Food for top predators and economically important species. Used by humans for food Pearls from oysters Shells used for jewelry