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Wiegand's Sedge Carex wiegandii Scientific Name Carex wiegandii Mackenzie Family Name Cyperaceae Sedge Family Did you know? Carex wiegandii is named in honor of Karl McKay Wiegand (1873-1942), a major botanist from New York (Fernald 1970). Wiegand worked at Cornell University and the herbarium at Cornell is known as the Wiegand Herbarium. Photo credits: Troy Weldy Summary Protection Endangered in New York State, not listed federally. This level of state protection means: listed species are those with: 1) 5 or fewer extant sites, or 2) fewer than 1,000 individuals, or 3) restricted to fewer than 4 U.S.G.S. 7 ½ minute topographical maps, or 4) species listed as endangered by U.S. Department of Interior. Rarity G4, S1 A global rarity rank of G4 means: This species is apparently secure globally (typically with more than 100+ populations), though it may be quite rare in parts of its range, especially at the periphery. A state rarity rank of S1 means: This plant is endangered/critically imperiled in New York because of extreme rarity (typically 5 or fewer populations or very few remaining individuals) or is extremely vulnerable to extirpation from New York due to biological factors. Conservation Status in New York There are four known sites and only one historical location. These sites are in remote Adirondack and Tug Hill peatlands that face few to no threats and are within well-protected landscapes. This is a difficult to identify and difficult to locate sedge that may be overlooked. Other bog areas within the Adirondacks and Tug Hill should be surveyed with this sedge in mind. NYNHP Conservation Guide - Wiegand's Sedge (Carex wiegandii) Short-term Trends Short term trends can not be assessed for populations in New York because there is not enough data available. Still, it is known that two populations have remained extant over the past twenty to thirty years. Long-term Trends There is very little historical data regarding C. wiegandii in New York. Therefore, long term trends are unknown. Conservation and Management Threats Populations in New York are not directly threatened. Some of the populations are very small and although this species generally occurs like this throughout its range, small populations are inherently vulnerable to even small scale changes in the environment such as flooding or natural disturbances. Conservation Strategies and Management Practices Currently no management is needed for populations of C. wiegandii in New York. Research Needs Specimens from two sites need to be found and/or collected. They should be verified. Further survey work should be done to assess the full extent of populations. Habitat In New York, Carex wiegandii occurs in acidic peatlands. It mostly occurs in the shade of conifers or alder thickets. It also is found in canopy openings and on the edge of open acidic peatlands. It occurs both on hummocks and depressions in these wetlands. The ground is usually carpeted with Sphagnum spp. (New York Natural Heritage Program 2006). It occurs in bogs, openings in acidic conifer, mixed, or alder swamps, and in wet acidic sandy or peaty meadows (Reznicek 2002). Sphagnum bogs and other wet, open places (Gleason and Cronquist 1991). Associated Ecological Communities Black Spruce-tamarack Bog A conifer forest that occurs on acidic peatlands in cool, poorly drained depressions. The characteristic trees are black spruce and tamarack; in any one stand, either tree may be dominant, or they may be codominant. Canopy cover is quite variable, ranging from open canopy woodlands with as little as 20% cover of evenly spaced canopy trees to closed canopy forests with 80 to 90% cover. Patterned Peatland A large peatland whose surface forms a gentle slope with a mosaic of high and low areas (relative to water levels). These high and low areas occur as narrow or broad bands of vegetation and pools that extend perpendicular to the direction of water flow across the NYNHP Conservation Guide - Wiegand's Sedge (Carex wiegandii) 2 slope of the peatland. Peat moss (Sphagnum) is the most abundant plant. Other Probable Associated Communities Highbush blueberry bog thicket Inland poor fen Pine barrens vernal pond Associated Species Speckled Alder (Alnus incana) Hoary Sedge (Carex canescens) Threeseeded Sedge (Carex trisperma var. trisperma) Tamarack (Larix laricina) Cinnamon Fern (Osmunda cinnamomea) Black Spruce (Picea mariana) Sphagnum spp. Northern Wild Raisin (Viburnum nudum var. cassinoides) Identification Comments Carex wiegandii is a grass-like tufted perennial. It has long strap-like leaves that are 1.7-5.0 mm wide. Numerous flowering/fruiting stems, 10-105 cm long, arise from the tufted leaves at the base of the plants. There are a few leaves on the stems and toward the tip of these stems are 4-6 flower/fruit clusters. These flower clusters have female flowers at the tip and male flowers at the base. The female flowers develop into fruits (perigynia) that are 2.6-3.8 mm long and 1.3-2.1 mm wide. Once mature the fruits easily fall from the plants (Reznicek 2002). Best Life Stage for Identifying This Species The easiest time to identify C. wiegandii is when it is in fruit but the fruit are not yet easily shedding. Leaves are critical for identification and ideally a full specimen in fruit is needed for a correct determination to be made. The Best Time to See Carex wiegandii is in immature to mature fruit from late June to early August although by late July the perigynia are starting to shed easily. Therefore, the best time to survey for this species is in the first two to three weeks of July. NYNHP Conservation Guide - Wiegand's Sedge (Carex wiegandii) 3 Fruiting Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec The time of year you would expect to find Wiegand's Sedge in New York. Similar Species Carex wiegandii is a very distinctive member of section Stellulatae. When robust it is unmistakable (Reznicek and Ball 1980). The wide leaves, widest 2.8-5.0 mm, separates it from most other closely related species in New York. Carex echinata in New York, usually has widest leaves up to 2.7 mm wide but can occasionally have leaves that are a little bit wider. Carex echinata has perigynium beaks (0.85-)0.95-2.0 mm long; perigynia beaks (0.40-)0.45-0.86 times the length of the body; and pistillate scales of wide leaved individuals 2.1-3.1 mm long (Reznicek 2002). Carex atlantica ssp. atlantica has widest leaves 1.6-4.0(-4.5) mm wide. It differs from C. wiegandii in having wider and wider to long perigynia. Carex atlantica ssp. atlantica has lower perigynia mostly 2.1-3.0 mm wide and mostly 1.1-1.7 times as long as wide vs. lower perigynia mostly 1.2-2.0 mm wide and mostly (1.5-)1.7-3.0 times as long as wide for C. wiegandii. Conservation Comments Carex wiegandii is a member of section Stellulatae. It was first named in 1931 by Mackenzie (Reznicek 2002). Taxonomy Kingdom Plantae Phylum Anthophyta Class Monocots (Monocotyledoneae) Order Cyperales Family Cyperaceae (Sedge Family) Additional Common Names Sedge Additional Resources NYNHP Conservation Guide - Wiegand's Sedge (Carex wiegandii) 4 Links Flora of North America http://efloras.org/florataxon.aspx?flora_id=1&taxon_id=242357629 New York Flora Atlas http://www.newyork.plantatlas.usf.edu/Plant.aspx?id=1140 USDA Plants Database http://plants.usda.gov/java/nameSearch?mode=sciname&keywordquery= CAREX+WIEGANDII NatureServe Explorer http://natureserve.org/explorer/servlet/NatureServe?searchName=CAREX+WIEGANDII Google Images http://images.google.com/images?q=CAREX+WIEGANDII Best Identification Reference Reznicek, A.A. 2002. Carex Linnaeus sect. Stellulate Kunth. Pages 326-331 in Flora of North America Editorial Committee (editors), Flora of North America, North of Mexico, Volume 23, Magnoliophyta: Commelinidae (in part): Cyperaceae. Oxford University Pre References Fernald, M.L. 1950. Gray's manual of botany. 8th edition. D. Van Nostrand, New York. 1632 pp. Gleason, Henry A. and A. Cronquist. 1991. Manual of Vascular Plants of Northeastern United States and Adjacent Canada. The New York Botanical Garden, Bronx, New York. 910 pp. Holmgren, Noel. 1998. The Illustrated Companion to Gleason and Cronquist's Manual. Illustrations of the Vascular Plants of Northeastern United States and Adjacent Canada. The New York Botanical Garden, Bronx, New York. Keys, Jr.,J.; Carpenter, C.; Hooks, S.; Koenig, F.; McNab, W.H.; Russell, W.;Smith, M.L. 1995. Ecological units of the eastern United States - first approximation (cd-rom), Atlanta, GA: U.S. Department of Agriculture, Forest Service. GIS coverage in ARCINFO format, selected imagery, and map unit tables. Mackenzie, K.K. 1931-1935. Cariceae. North American Flora 18: 1-478. NatureServe. 2005. NatureServe Central Databases. Arlington, Virginia. USA New York Natural Heritage Program. 2010. Biotics database. New York Natural Heritage Program. New York State Department of Environmental Conservation. Albany, NY. Reschke, Carol. 1990. Ecological communities of New York State. New York Natural Heritage Program, New York State Department of Environmental Conservation. Latham, NY. 96 pp. plus xi. Reznicek, A.A. and P.W. Ball. 1980. The taxonomy of Carex section Stellulatae in North America north of Mexico. Contr . Unw. Mich. Herb. 14:153-203. Rhoads, A.F., and W.M. Klein, Jr. 1993. The vascular flora of Pennsylvania: Annotated checklist and atlas. American Philosophical Society, Philadelphia, PA. 636 pp. Rothrock, P.E. 1978. Carex wiegandii Mackenzie (Cyperaceae) New for Pennsylvania. Rhodora 80:451. A78ROT01PAUS. Weldy, T. and D. Werier. 2010. New York flora atlas. [S.M. Landry, K.N. Campbell, and L.D. Mabe NYNHP Conservation Guide - Wiegand's Sedge (Carex wiegandii) 5 (original application development), Florida Center for Community Design and Research http://www.fccdr.usf.edu/. University of South Florida http://www.usf.edu/ New York Natural Heritage Program This project is made possible with funding from: 625 Broadway, 5th Floor, - New York State Department of Environmental Conservation Hudson River Albany, NY 12233-4757 Estuary Program Phone: (518) 402-8935 - Division of Lands & Forests, Department of Environmental Conservation [email protected] - New York State Office of Parks, Recreation and Historic Preservation Information for this guide was last updated on Oct 28, 2015 This guide was authored by NYNHP Conservation Guide - Wiegand's Sedge (Carex wiegandii) 6