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Phylum Cnidaria General Information: • Cnidaria means ‘a stinging thread’ • Examples: sea anemones, corals, jellyfish, freshwater Hydra, and the Portuguese manof-war • Habitat: marine and fresh water Soft Coral Sea Anemone, Clover Point, Victoria B.C. Portuguese Man Of War Brain Coral Corals are an important part of BC marine ecosystems! •In BC, deep water corals such as the red tree or “Gorgonian coral” provide shelter for juvenile rockfish such as the yelloweye rockfish or sharpchin rockfish Characteristics: • All cnidarians have a basic radial symmetry and possess only 2 layers of living tissues. • 2 tissue layers: ectoderm and endoderm • Contain a mesoglea • Some possess an exoskeleton (for protection) made up of CaCO3 (e.g. corals) Characteristics • Have 2 body forms: a. Polyp (sessile) – e.g. hydra, coral, or sea anemone b. Medusa (free swimming) – bell shaped – e.g. jellyfish • Some cnidarians contain only one body shape during their life cycle, while others contain both during their life cycle • When a cnidarian exhibits the 2 body forms in its life cycle, it is called polymorphism (‘many shapes’) • The evolution of a primitive nervous system sets these organisms apart from the sponges – called a nerve net • They contain nematocysts found on the tentacles used for capturing prey and for protection • Mesoglea is sandwiched between ectoderm and endoderm • There are amoebocytes in the mesoglea of cnidarians, much like in sponges • Cnidarians possess a simple digestive system – the gastrovascular cavity – where digestion and circulation of nutrients occur http://shapeoflife.org/video/cnidarians-lifemove Sea Anemones – Class Anthozoa: • • • • No medusa stage They are all marine Polyp body form exclusively Reproduce asexually through longitudinal or transverse fission (like binary fission, but on a much larger scale) • Are primarily carnivorous – capture food using nematocyst-studded tentacles and transfer it to a central mouth opening • There are around 6000 species of sea anemones Coral – Class Anthozoa • Colonial form of Class Anthozoa • Can have soft and hard corals • Hard corals secrete external CaCO3 skeletons and are called ‘stony corals’ • Stony corals make up coral reefs (not all make reefs though) • Coral add about 10 kg of additional CaCO3 per m2 of reef yearly • Soft corals do not have hard external skeletons The Anatomy of A Coral • Many corals have symbiotic green algae called zooxanthellae living in their polyps • Warmer ocean temperatures can cause these algae to die, resulting in a phenomenon known as “coral bleaching” Jelly Fish – Class Scyphozoa • Only a few hundred species – nearly all are marine • The body is in the form of an inverted cup, with nematocyst-studded tentacles extending downward from the cup. • Some jellyfish have symbiotic unicellular algae that give them nutrients • The life cycle of jelly fish involves the two different body forms: polyp and medusa A Typical Jellyfish Life Cycle A Typical Cnidarian - Hydra Structure: • Polyp body form Locomotion: • Sessile; however, some can somersault and glide along while secreting mucus Ingestion, Digestion, Elimination • Food is captured by nematocysts in the tentacles by stinging or stunning prey • Tentacles push food through mouth into gastrovascular cavity • Flagella line the endoderm layer and create a current • The endoderm cells secrete enzymes to break down food • Food is taken into cells and nutrients are circulated by amoeboid cells • Wastes are eliminated out through the mouth The Nematocyst Nematocysts vary in shape, but all are adapted for stinging or stunning prey In the sea anemone on the right, nematocyst aggregations are visible as the dark pink lines Respiration • O2 enters the cells by diffusion • CO2 leaves the cells by diffusion Nervous System • Have a nerve net within the mesoglea • When stimulated, the nerve net will respond by contracting / relaxing the muscles • Have sensory cells that are sensitive to touch, light, chemicals, and balance Muscular System • Latitudinal and circular muscle fibres are found in the endo / ectoderm layers Excretion • Nitrogenous wastes leave the cells by diffusion • Wastes enter the gastrovasuclar cavity and leave via the mouth Reproductive System A. Asexual • Budding (bud breaks away) • Regeneration (re-growth of missing parts) A Hydra Budding Reproductive System B. Sexual • Some are hermaphrodites • Ovaries and testes grow on the side of adults • Ovary produces an egg and remains attached to the adult • Testes produce the sperm which swim to the egg, resulting in a zygote • Zygote develops into a ball of cells called the blastula, which develops into a ciliated larva called a planula, which can swim to another location, settle, and develop into a polyp adult Ecological Importance of Cnidarians • Are extremely important in the ecology of tropical and temperate oceans in the form of coral (eg Great Barrier Reef) • Corals provide habitats and food for many species of fish and invertebrates • Reefs protect the land from ocean waves • Cnidarians form symbiotic relationships with other organisms living among the tentacles (clown fish and sea anemones) The Importance of Deepwater Corals •Throughout the world, many deepwater coral reefs are being destroyed by trawlers, removing vital nursery grounds for many fish species Cnidarians and Medicine • Used in medical research for their chemicals (cancer therapy) • Stony corals are now being used in certain surgical procedures. Because they have many pores for blood vessels to grow through, small pieces of coral are now being used in bone grafts, particularly in face and jaw reconstruction, and in arm and leg surgery • These pieces are eventually replaced by normal bone tissue Evolutionary Significance • Cnidarians are the first animals to have organized true cell tissues • They are therefore the first true multicellular organisms to appear • They are the first organisms to have muscle tissue and nervous tissue • Cnidarians, having radial symmetry are thought to share a common ancestor with all other multicellular animals