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Transcript
Chapter 53
• Population ecology is the study of
populations in relation to environment,
including environmental influences on
density and distribution, age structure,
and population size
Population Ecology
Concept 53.1: Dynamic biological
processes influence population
density, dispersion, and
demographics
• A population
is a group of individuals of
a single species living in the same
general area
Density and Dispersion
• Density is the number of individuals per
unit area or volume
• Dispersion is the pattern of spacing
among individuals within the boundaries
of the population
Density: A Dynamic Perspective
• In most cases, it is impractical or
impossible to count all individuals in a
population
• Sampling techniques can be used to
estimate densities and total population
sizes
• Population size can be estimated by
either extrapolation from small samples,
an index of population size, or the markrecapture method
• Density is the result of an interplay
between processes that add individuals
to a population and those that remove
individuals
• Immigration is the influx of new
individuals from other areas
• Emigration is the movement of
individuals out of a population
1
Fig. 53-3
Demographics
Births
Deaths
Births and immigration
add individuals to
a population.
Immigration
Deaths and emigration
remove individuals
from a population.
• Demography is the study of the vital
statistics of a population and how they
change over time
• Death rates and birth rates are of
particular interest to demographers
Emigration
Fig. 53-5
Survivorship Curves
• A survivorship curve is a graphic way of
representing the data in a life table
• The survivorship curve for Belding’s
ground squirrels shows a relatively
constant death rate
Number of survivors (log scale)
1,000
100
Females
10
Males
1
0
2
8
4
6
Age (years)
10
• Survivorship curves can be classified into
three general types:
– Type I: low death rates during early and middle
life, then an increase among older age groups
– Type II: the death rate is constant over the
organism’s life span
– Type III: high death rates for the young, then a
slower death rate for survivors
Number of survivors (log scale)
Fig. 53-6
1,000
I
100
II
10
III
1
0
50
Percentage of maximum life span
100
2
Reproductive Rates
• For species with sexual reproduction,
demographers often concentrate on
females in a population
• A reproductive table, or fertility
schedule, is an age-specific summary of
the reproductive rates in a population
• It describes reproductive patterns of a
population
• Some plants produce a large number of
small seeds, ensuring that at least some
of them will grow and eventually
reproduce
• In animals, parental care of smaller
broods may facilitate survival of offspring
• Zero population growth occurs when
the birth rate equals the death rate
• Most ecologists use differential calculus
to express population growth as growth
rate at a particular instant in time:
∆N =
rN
∆t
where N = population size, t = time, and r = per
capita rate of increase = birth – death
• Exponential population growth is
population increase under idealized
conditions
• Under these conditions, the rate of
reproduction is at its maximum, called the
intrinsic rate of increase
Fig. 53-10
2,000
dN
= 1.0N
dt
Population size (N)
Exponential Growth
1,500
dN
= 0.5N
dt
1,000
500
0
0
5
10
Number of generations
15
3
Fig. 53-11
Concept 53.4: The logistic model describes how a population
grows more slowly as it nears its carrying capacity
Elephant population
8,000
6,000
4,000
2,000
0
1900
1920
1940
Year
1960
1980
The Logistic Model and Real
Populations
Fig. 53-12
Exponential
growth
Population size (N)
2,000
dN
= 1.0N
dt
1,500
• The growth of laboratory populations of
paramecia fits an S-shaped curve
• These organisms are grown in a constant
environment lacking predators and
competitors
K = 1,500
Logistic growth
1,000
• Exponential growth cannot be sustained
for long in any population
• A more realistic population model limits
growth by incorporating carrying capacity
• Carrying capacity (K) is the maximum
population size the environment can
support
dN
= 1.0N
dt
1,500 – N
1,500
500
0
0
5
10
Number of generations
15
Population Dynamics
Number of Daphnia/50 mL
Number of Paramecium/mL
Fig. 53-13
1,000
800
600
400
200
0
• The study of population dynamics
focuses on the complex interactions
between biotic and abiotic factors that
cause variation in population size
180
150
120
90
60
30
0
0
5
10
Time (days)
15
(a) A Paramecium population in the lab
0
20
40
60
80 100 120
Time (days)
140
160
(b) A Daphnia population in the lab
4
Fig. 53-18
Stability and Fluctuation
2,100
Number of sheep
1,900
• Long-term population studies have
challenged the hypothesis that
populations of large mammals are
relatively stable over time
• Weather can affect population size over
time
1,700
1,500
1,300
1,100
900
700
500
0
1955
1965
1975
1985
Year
1995
2005
Fig. 53-19
2,500
50
40
2,000
30
1,500
20
1,000
10
500
0
1955
1975
1985
Year
0
2005
1995
Fig. 53-20
160
Snowshoe hare
120
9
Lynx
80
6
40
3
Number of lynx
(thousands)
• Some populations undergo regular boomand-bust cycles
• Lynx populations follow the 10 year
boom-and-bust cycle of hare populations
• Three hypotheses have been proposed to
explain the hare’s 10-year interval
1965
Number of hares
(thousands)
Population Cycles: Scientific
Inquiry
Moose
Number of moose
Wolves
Number of wolves
• Changes in predation pressure can drive
population fluctuations
0
0
1850
1875
1900
Year
1925
5
The Global Human Population
Fig. 53-22
• The human population increased
relatively slowly until about 1650 and then
began to grow exponentially
6
5
4
3
2
The Plague
1
Human population (billions)
7
0
8000
B.C.E.
4000 3000 2000 1000
B.C.E. B.C.E. B.C.E. B.C.E.
0
1000
C.E.
2000
C.E.
Fig. 53-23
2.2
2.0
1.8
Annual percent increase
• Though the global population is still
growing, the rate of growth began to slow
during the 1960s
1.6
1.4
2005
1.2
Projected
data
1.0
0.8
0.6
0.4
0.2
0
1950
Regional Patterns of Population
Change
• To maintain population stability, a
regional human population can exist in
one of two configurations:
– Zero population growth =
High birth rate – High death rate
– Zero population growth =
Low birth rate – Low death rate
• The demographic transition is the move
from the first state toward the second
state
1975
2000
Year
2025
2050
Limits on Human Population Size
• The ecological footprint concept
summarizes the aggregate land and
water area needed to sustain the people
of a nation
• It is one measure of how close we are to
the carrying capacity of Earth
• Countries vary greatly in footprint size
and available ecological capacity
6
Fig. 53-27
• Our carrying capacity could potentially be
limited by food, space, nonrenewable
resources, or buildup of wastes
Log (g carbon/year)
13.4
9.8
5.8
Not analyzed
Food From the Sea
• What types of organisms are harvested?
Worldwide
Marine Catch
and Mariculture
– Finfish (about 90% of worldwide harvest)
– Shellfish
– Other species such as jellyfish, sea
cucumbers, polychaetes and seaweed
– While seafood represents only about 1% of
the food consumed each year, it represents
about 30% of total animal protein consumed
Atlantic bluefin tuna Thunnus
thynnus
• Can grow >300 cm; 680 kg
• Extremely streamlined, one of the ocean’s
fastest swimmers, endothermic
Bluefin as food
• 2001 440 pound
tuna sold for
$220,000
($500/pound)
• Farm in oceanic
pens
• Spotter planes
and electric
harpoons
7
Optimal Yield and Overfishing
• SeaSea-life species are renewable resources
• However, for a fishery to last longlong-term, it
must be fished in a sustainable way
• The sustainable yield is the amount that
can be caught and just maintain a
constant population size
Collapse of a Fishery
• A fishery is regarded as collapsed if
numbers fall to 10% of historic highs
• It is estimated that oneone-third of fisheries
are already collapsed
• A 2006 study indicates that all major
fisheries will collapse by 2050 if protective
measure are not taken to better manage
and protect these resources
Managing the Resources
• Management can be difficult for many
reasons:
– Maximum sustainable yield is difficult to
calculate
– Harvested species may compete with other
species and fishing pressure may affect
competitive balance
– Real fisheries are more complex than models
– High seas are “common property”
property”
• Bluefin tuna harpoon
• http://www.youtube.com/watch?v=tL1te9SbLs&feature=related
• crab pot
• http://www.youtube.com/watch?v=Zd_OP
FfpRdk
• tuna farming
• http://www.youtube.com/watch?v=XIbGTw
LGZNU&feature=related
8