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Visualizing Environmental Science
Ecosystems and Evolution
Chapter 6
Copyright © 2014 John Wiley & Sons, Inc. All rights reserved.
Earth’s Major Biomes
• Biome—a large, relatively
distinct terrestrial region
with similar climate (longterm weather pattern), soil,
plants, and animals,
regardless of where it
occurs in the world
– Encompasses many
interacting ecosystems
© 2014 John Wiley & Sons, Inc. All rights reserved.
Factors That Affect Biomes
• Temperature and precipitation
have a predominant effect on
biome distribution
– Latitude
• Temperature most important factor
near poles
• Precipitation more important in
tropical and temperate regions
• Elevation also affects biomes
– Going from warmer to colder
climates
© 2014 John Wiley & Sons, Inc. All rights reserved.
Arctic Tundra
• Treeless biome in the far north
that consists of boggy plains
covered by lichens and mosses
• Permafrost—layer of permanently
frozen ground
– Climate change causing permafrost to
thaw
• Harsh, cold winters and extremely
short summers and seasonal snow
melting
• Alpine tundra occurs at higher
elevations of mountains, above
tree line
• Low primary productivity and low
resilience
© 2014 John Wiley & Sons, Inc. All rights reserved.
Boreal Forest/Taiga
• Coniferous forests of pine,
spruce, and fir, in the Northern
Hemisphere, just south of tundra
• Winters extremely cold and
severe
• Little precipitation, 20 in/year
• Soil is acidic and mineral poor
• Caribou, wolves, bear, moose,
rodents, rabbits, lynx, birds in
summer, lots of insects
• Top source of industrial wood
and fiber, leading to loss of forest
© 2014 John Wiley & Sons, Inc. All rights reserved.
Temperate Rain Forest
• Coniferous forest with high
precipitation, dense fog and cool
weather
• Northwest coast of NA, SE Australia, SA
• Mild winters, cool summers
• Slow decay, poor soil
• Hemlock, fir, cedar, spruce, epiphytes,
mosses, lichens and ferns
• Squirrels, wood rats, elk, mule deer,
birds, amphibians and reptiles
• High producer of lumber and pulpwood
•
Overharvesting old-growth forest is an
issue; see chapter 13 for debate over clearcutting
© 2014 John Wiley & Sons, Inc. All rights reserved.
Temperate Deciduous Forest
• Forest biome that occurs in
temperate areas where
precipitation ranges from 30–50
in/year
• Hot summers, cold winters
• Topsoil rich in organic matter
• Broad-leaf hardwood trees, lose
leaves seasonally
• Originally puma, wolves, and
bison, now absent
• Deer, bears, small mammals,
birds
• Original forests in Europe and NA
mostly destroyed by logging and
urbanization
© 2014 John Wiley & Sons, Inc. All rights reserved.
Temperate Grassland
• Tallgrass prairies and shortgrass
prairies
• Hot summers, cold winters, and less
rainfall (10 – 30 in/year) than in
temperate deciduous forest biome
• Soil rich in organic matter
• More than 90% of tallgrass prairies
were plowed for agriculture, NA’s
rarest biome
• Bison graze on mixed-grass prairie in
Custer State Park, South Dakota
© 2014 John Wiley & Sons, Inc. All rights reserved.
Chaparral
• Mild, moist winters; hot, dry
summers
• Referred to as having a
“Mediterranean climate”
• Small-leaved evergreen shrubs
and small trees dominate
• Lush vegetation during rainy
winter season
• Wildfires common in late
summer and autumn
– Many fire adapted plants
that grow after a fire
• Thin, unfertile soil
© 2014 John Wiley & Sons, Inc. All rights reserved.
EnviroDiscovery
Using Goats to Fight Fire
• California has 6000 wildfires
each year with many people
living in fire-prone chaparral
• Goats are being used to clear
hills of vegetation during the 6month fire season
– 350 goats can clear an acre of heavy
brush per day
• Rare/endangered plant species
are fenced off
© 2014 John Wiley & Sons, Inc. All rights reserved.
Desert
•
•
•
•
•
•
•
Plant growth limited by lack of
precipitation
Found in both temperate and tropical
regions
Daily temperature extremes
Less than 10 inches of rain/year
Sparse vegetation that includes cacti,
yucca, and sagebrush
Soil is low in organic matter but high in
mineral content
Animals
– Typically small, mostly nocturnal
– Insects, arachnids, desert-adapted
amphibians, many reptiles
•
Threatened by human encroachment and
environmental damage from off-road
vehicles
– Soil easily eroded and less vegetation grows to
support native animals
© 2014 John Wiley & Sons, Inc. All rights reserved.
Savanna
• Tropical grassland
• Large herds of herbivores such as
wildebeest, antelope, giraffe, zebra,
elephant
• Large carnivores such as lions and
hyenas
• Low or intense seasonal rainfall, 34–60
in/year, but with prolonged dry periods
• Widely scattered or clumped trees
such as the acacia
• Low mineral-content soil
• Savanna lost as land is converted into
rangeland for cattle
• Africa, N Australia, South America, W
India
© 2014 John Wiley & Sons, Inc. All rights reserved.
Tropical Rain Forest
• Lush, species rich forest that occurs
where the climate is warm and
moist year-round
• Rains almost daily, 80–180 in/year
• Weathered, mineral-poor soil
• High species richness and diversity
• Three layers of vegetation
• In tropical dry forests precipitation
seasonal
• Industrial expansion and human
population growth threaten the rain
forests
© 2014 John Wiley & Sons, Inc. All rights reserved.
Florida’s Terrestrial Habitats
• Imperceptible elevation changes
• Pine Flatwoods
– Acidic soil
– Clay hardpan
• Stores water
• Lessens root
penetration
– Fire dependent
– Species
• Saw palmetto
• Slash pine
• Deer, fox, raccoon, possum, panther, snakes, burrowing owl
Image and Habitat information from the South Florida Water Management District www.sfwmd.gov
Florida’s Terrestrial Habitats
• Scrub
– Highest elevation
– Sandy, well drained
soil
– Species
• Sand Pine
• Scrub oak
• Deer, fox, raccoon, possum, panther, snakes, gopher tortoise,
bobcat
Image and Habitat information from the South Florida Water Management District www.sfwmd.gov
Florida’s Terrestrial Habitats
• Hardwood Hammock
– Lowest elevation
– Cooler
– Species
•
•
•
•
Gumbo Limbo
Oaks
Cabbage Palm (state tree)
Deer, fox, raccoon, possum, panther, snakes,
Image and Habitat information from the South Florida Water Management District www.sfwmd.gov
Aquatic Ecosystems
• Aquatic ecology
– Freshwater ecosystems include
• Standing-water (lakes and
ponds)
• Flowing-water (rivers and
streams)
• Wetlands (marshes and swamps)
– Categories of organisms
• Plankton—phytoplankton and
zooplankton
• Nekton—fish and turtles
• Benthos—bottom-dwellers
© 2014 John Wiley & Sons, Inc. All rights reserved.
Freshwater Ecosystems
• Occupy 2% of earth’s surface, yet play important role in
hydrologic cycle
• Standing-water, lakes and ponds have three zones
– Littoral, limnetic, profundal
• Zonation accentuates thermal stratification, temperature
changes with depth
• Human effects include eutrophication, unnatural nutrient
enrichment
© 2014 John Wiley & Sons, Inc. All rights reserved.
Freshwater Wetlands
• Marshes, dominated by grass-like
plants, and swamps, dominated by
woody shrubs
• Shallow fresh water cover for at least
part of the year
• Water-tolerant vegetation
• Waterlogged soils
– Anaerobic, low rate of decomposition
– Rich in organic material
• Ecosystem services include wildlife
habitat, flood mitigation and filtration of
ground water
• Wetlands threatened by pollution,
development and agriculture
© 2014 John Wiley & Sons, Inc. All rights reserved.
Brackish Ecosystems: Estuaries
• Estuary—coastal body of water,
partly surrounded by land, with
access to the ocean and fresh water
from a river
• Water levels rise and fall with tides
• Salinity fluctuates from fresh water
to brackish
• Highly productive ecosystems with
rapid nutrient circulation
• Salt marshes—shallow wetlands with
salt-tolerant grasses
• Mangrove forests—tropical
equivalent of salt marshes
© 2014 John Wiley & Sons, Inc. All rights reserved.
Three Species of Florida Mangroves
•Red Mangroves usually grow near the shore
of the water. It has red roots and is often
referred to as the “walking mangrove”
because its roots raise over the water.
•Black Mangroves grow in higher areas Their
roots spread near the trunk in shapes of
fingers or pencils pointing up, exposing the
roots to needed oxygen.
•White Mangroves grow even higher than the
black mangrove, and its roots aren’t visible.
These trees tend to get rid of salt on the
backs of the leaves.
•Significant losses due to coastal
development and aquaculture
Community Responses to Changing Conditions
Over Time: Succession
• Ecological succession—the process of community development over time,
through a sequence of species
• Resident species modify the environment, making it more suitable for later
species
• Former concept of a stable ‘climax community’ has been replaced with reality
that communities continue to change
• Primary and secondary succession have specific species compositions
© 2014 John Wiley & Sons, Inc. All rights reserved.
Primary Succession
• Change of species over time in a previously uninhabited
environment
• No soil, bare rock surfaces
• Begins with pioneer community composed of acidic lichens that
break up rock and form soil
• Lichensmossesshrubseventually specific trees
• Note the stages of primary succession on glacial moraine
© 2014 John Wiley & Sons, Inc. All rights reserved.
Primary Succession
• Sandbars
• Lava flows that have cooled
© 2014 John Wiley & Sons, Inc. All rights reserved.
Secondary Succession
• Change in species composition after a
disturbance in an area previously
inhabited
• Abandoned farmland, forest fire,
clear-cut forest
• Soil already present
• Can take more than
100 years for secondary
succession to occur
• Typically crabgrasshorseweedpine treeshardwood trees
© 2014 John Wiley & Sons, Inc. All rights reserved.
Developing and Understanding of Evolution
© 2014 John Wiley & Sons, Inc. All rights reserved.
Galápagos Islands
• Evolution—the cumulative
genetic changes in populations
that occur during successive
generations
• Charles Darwin (1809–1882)
proposed the mechanism of
evolution (i.e., not idea of
evolution) in “The Origin of
Species by Means of Natural
Selection” (1859)
– Environment plays crucial role in
which traits are inherited
– Accumulation of favorable traits
leads to increased survival (fitness)
Evolution Through Natural Selection
• Four observations guide natural
selection (NS)
–
–
–
–
High reproductive capacity
Limits on population growth
Heritable variation
Differential reproductive success
© 2014 John Wiley & Sons, Inc. All rights reserved.
Darwin’s Finches
• While on the Galapagos Islands
off the coast of Ecuador,
Darwin studied plants and
animals on each island,
including 14 species of finches
• He concluded that the finches
had a common mainland
ancestor, but had become
geographically isolated and
adapted to different diets
© 2014 John Wiley & Sons, Inc. All rights reserved.
Environmental InSight
Evidence for Evolution
• Fossil record
– Fossils show how organisms
evolved over time
• Comparative anatomy
– Similarities among organisms
demonstrate how they are
related
• Biogeography
– The study of geographic
locations of organisms
• Molecular biology
– Showing relationships on a
molecular level such as
sharing an enzyme or
nucleotide base
© 2014 John Wiley & Sons, Inc. All rights reserved.