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2/11/2015 Fermentation - Chemwiki Sign In Forgot Password Register username username password password Sign In If you like us, please share us on social media. The latest UCD Hyperlibrary newsletter is now complete, check it out. ChemWiki BioWiki Periodic Table of the Elements GeoWiki Reference Tables StatWiki Physical Constants PhysWiki MathWiki SolarWiki Units & Conversions Lab Techniques ChemWiki: The Dynamic Chemistry E-textbook > Biological Chemistry > Metabolism > Glycolysis > Fermentation Fermentation Fermentation is the process by which living organisms recycle in the absence of oxygen. is a required molecule necessary for the oxidation of Glyceraldehyde-3-phosphate to produce the high energy molecule 1,3-bisphosphoglycerate (Step 6 of Glycolysis). Fermentation occurs in the cytosol of cells. Introduction Because is used in Glycolysis it is important that living cells have a way of recycling from . One way that a cell recycles is by reducing oxygen in the electron transport chain. As transfers its electrons to oxygen in the form of a hydride ion it is reduced to which can then be used again in step 6 of Glycolysis. Another way that is recycled from is by a process called fermentation. Lactic acid fermentation in contracting muscle Lactic acid fermentation occurs by converting pyruvate into lactate using the enzyme Lactate dehydrogenase and producing in the process. This process takes place in oxygen depleted muscle and some bacteria. It is responsible for the sour taste of sauerkraut and yogurt. is required for the oxidation of glyceraldehyde-3-P to produce 1,3-Bisphosphoglycerate (Step 6 of Gycolysis). If the supply of is not replenished by the ETC or fermentation glycolysis is unable to proceed. Fermentation is a necessary process for anaerobic organisms to produce energy. The yield of energy is much less than if the organism were to continue on through the TCA cycle and ETC, but energy is produce nonetheless. Alcoholic fermentation in yeast The purpose of fermentation in yeast is the same as that in muscle and bacteria, to replenish the supply of NAD+ for glycolysis, but this process occurs in two steps: 1. Alcoholic fermentation consists of pyruvate being first converted into acetaldehyde by the enzyme pyruvate decarboxylase and releasing . 2. In the second step acetaldehyde is converted into ethanol using alcohol dehydrogenase and producing in the process. It is this recycled that can be used to continue on with glycolysis. References http://chemwiki.ucdavis.edu/Biological_Chemistry/Metabolism/Glycolysis/Fermentation 1/2 2/11/2015 Fermentation - Chemwiki Garrett, H., Reginald and Charles Grisham. Biochemistry. Boston: Twayne Publishers, 2008. Raven, Peter. Biology. Boston: Twayne Publishers, 2005. Problems 1. 2. 3. 4. 5. Draw the chemical structures of pyruvate, ethanol and lactate (the reactant and products of fermentation) Why is fermentation necessary? (Hint: see step 6 of Glycolysis) What type of environment is necessary for fermentation to occur? Where does fermentation occur? What part of the cell? Explain the alternative to fermentation and why it is able to proceede. (Hint: Final electron acceptor) Contributors Darik Benson (Undergraduate University California Davis) Mike Blaber (Florida State University) © Copyright 2015 Chemwiki Powered by MindTouch ® Unless otherwise noted, content in the UC Davis ChemWiki is licensed under a Creative Commons Attribution-Noncommercial-Share Alike 3.0 United States License. Permissions beyond the scope of this license may be available at [email protected]. Questions and concerns can be directed toward Prof. Delmar Larsen ([email protected]), Founder and Director. Terms of Use http://chemwiki.ucdavis.edu/Biological_Chemistry/Metabolism/Glycolysis/Fermentation 2/2