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Photosynthesis:
(Note: subscripts will not appear correctly in the html version of the notes below)
Ultimately, almost all life is dependent on plants (or algae, etc.):
- plants take sunlight and convert it into energy (sugar molecules).
sunlight
6 CO2 + H2O -----------------------> C6H12O6 + 6O2
Before we tackle photosynthesis, some background:
- autotroph - means “self feeder” Plants get their food from sunlight (above
equation).
Within plants (and eukaryotic algae), we have chloroplasts. These are the parts of
a plant that are green and actually carry out photosynthesis [OVERHEAD, fig.
7.2, p. 109].
- within the chloroplast there are membranous sacs called thylakoids.
- the fluid between the thylakoids and the inside membrane is known as
the stroma.
Two stages to photosynthesis [OVERHEAD, fig. 7.5, p. 111]:
- light reactions - take place within the thylakoid membranes.
- convert light energy into chemical energy. Also produce O2 as a waste
product.
- this energy is stored as ATP and NADPH. NADPH stores energy in by
attaching to electrons. Remember this from respiration (NADPH is
similar to NADH).
- Calvin cycle - takes place within the stroma.
- this is where CO2 is converted into sugars, using the energy provided by
the light reactions.
- does not require light, though usually only active if light reactions are
going.
Source of energy:
- electromagnetic radiation (light). All light has energy in it (this should b
obvious).
- Chlorophyll can absorb visible light, particularly in the blue-violet ranges, and
then the red ranges.
- light that is reflected is not used. Green is reflected, so plants look green.
- see [OVERHEAD, fig. 7.6B, p. 112].
- other pigments (carotenes, xanthophyll) can help with this. These are the
pigments that become visible when leaves change color (chlorophyll falls
apart quickly, and what we see is these other pigments).
Photons and electrons:
- light waves consist (glossing over a lot of physics here) in part of photons.
- these photons hit the chlorophyll, and cause the chlorophyll to donate a high
energy electron to other molecules.
- if interested, the details are in section 7.7, p. 113.
Using electrons [OVERHEAD, fig. 7.8, p. 116]:
- the high energy electrons are used as follows (these processes technically use
electron transport chains):
- there exist two photosystems. Each uses light to put energy into
electrons.
- photosystem II donates electrons to photosystem I. In the process, it uses
up water and makes O2 (it replaces it’s electrons from water) and
Hydrogen ions.
- photosystem I gets its electrons from photosystem II (via an electron
transport chain), then donates its high energy electrons to an electron
transport chain that makes NADPH.
- further, the Hydrogen ions that are released with the Oxygen is used to
convert (“phosphorylate) ADP into ATP (a net increase in energy).
- how do you convert H+ ions into ATP? We'll discuss that during
cellular respiration.
- end result is ATP and NADPH are made from the light reactions.
Calvin cycle:
- No details here (if interested, see p. 118).
- The Calvin cycle uses the energy made in the light reactions (ATP & NADPH)
to convert CO2 into a sugar called G3P.
- G3P can easily be converted into familiar sugars such as glucose.
- Essentially, one molecule of CO2 is used at a time. Thus to make glucose (a 6
carbon sugar, we need to go through the cycle 6 times.
Energy usage:
The sugars made through these processes power everything that the plant needs
(incl. cellular respiration & the Krebs cycle), as well as all animal life on the
planet (eating meat only means eating something that ultimately eats plants if one
goes down the food chain far enough).
Finally, a little bit about C4 plants:
The process we’ve looked at requires CO2 to enter the leaf, and O2 to leave.
On hot dry days, stoma (openings in leaves & stems that let air in or out) will
close (to prevent water loss), preventing photosynthesis.
C4 and CAM plants have adapted to dry climates by being able to utilize much
lower concentrations of CO2 than C3 plants. Thus, keeping their stoma closed
doesn’t hurt photosynthesis as much.
- Way too many details (if you’re interested) are on p. 118.