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Animal Diversity I: Invertebrates Copyright © 2005 Pearson Prentice Hall, Inc. Animal Key Features Copyright © 2005 Pearson Prentice Hall, Inc. Anatomical Branch Points on the Animal Evolutionary Tree • An evolutionary tree of some major animal phyla (F22.1 p. 424) Copyright © 2005 Pearson Prentice Hall, Inc. ecdysozoans Cnidaria Porifera (sponges) (jellyfish, corals, anemones) Ctenophora (comb jellies) bilateral protostomes lophotrochozoans Nematoda Arthropoda (roundworms) (insects, arachnids, crustaceans) Platyhelminthes (flatworms) Annelida (segmented worms) deuterostomes Mollusca (clams, snails, octopods) Echinodermata (sea stars, sea urchins) cuticle molted protostome development radial symmetry no tissues tissues Sponges: Lack Tissues deuterostome development bilateral symmetry Radial or Bilateral Symmetry Chordata (lancelets, vertebrates) Anatomical Branch Points on the Animal Evolutionary Tree •Animals with Tissues Exhibit Either Radial or Bilateral Symmetry Body symmetry and cephalization (F22.2 p. 425) Radial symmetry Bilateral symmetry central axis plane of symmetry anterior plane of symmetry posterior Anatomical Branch Points on the Animal Evolutionary Tree – Embryonic Tissue Layers • Radially Symmetrical Animals Have Two • Bilaterally Symmetrical Animals Have Three –Bilateral Animals Have Heads – Most Bilateral Animals Have Body Cavities – Body Cavity Structure Varies Among Phyla • Body cavities Copyright © 2005 Pearson Prentice Hall, Inc. (F22.3 p. 426) Anatomical Branch Points on the Animal Evolutionary Tree body wall } coelom } digestive tract digestive cavity “True” coelom (annelids, chordates) Body cavities body wall } pseudocoelom body } wall digestive tract digestive cavity “False” or pseudocoelom (roundworms) digestive cavity digestive tract No coelom (cnidarians, flatworms) (F22.3 p. 426) Anatomical Branch Points on the Animal Evolutionary Tree • Simpler Body Cavities Evolved from Coelomate Body Plans • Bilateral Organisms Develop in One of Two Ways – Protostomes Deuterostomes • Protostomes: Two Distinct Evolutionary Lines – Ecdysozoans Copyright © 2005 Pearson Prentice Hall, Inc. Lophotrochozoans Comparison of the Major Animal Phyla (T22.1 p. 428) osculum epithelial cell pore Major Animal Phyla spicules (water flow) amoeboid cell pore cell collar cell Sponges: Simple Body Plan (F22.4 p. 428) (water flow) Sponge diversity (F22.5 p. 429) Major Animal Phyla • Cnidarians: Well-Armed Predators – Cnidarian diversity (F22.6 p. 430) – Polyp and medusa (F22.7 p. 431) – Cnidarian weaponry: the cnidocyte (F22.8 p. 431) Copyright © 2005 Pearson Prentice Hall, Inc. Major Animal Phyla Cnidarians: WellArmed Predators Cnidarian diversity (F22.6 p. 430) Major Animal Phyla Polyp mouth lining of gastrovascular cavity Medusa lining of gastrovascular cavity body wall tentacle gastrovascular cavity column body wall mesoglea mesoglea mouth foot gastrovascular cavity Polyp and medusa (F22.7 p. 431) tentacle trigger filament trigger nuclei Cnidarian weaponry: the cnidocyte (F22.8 p. 431) Major Animal Phyla • Flatworms – Organs – Lack Respiratory & Circulatory Systems – Flatworm organ systems (F22.9 p. 432) – The life cycle of the human pork tapeworm (F22.10 p. 433) Copyright © 2005 Pearson Prentice Hall, Inc. Major Animal Phyla Digestive system eyespots gastrovascular cavity Excretory system Nervous system brain excretory pore pharynx (with mouth at tip) nerve cord Flatworm organ systems (F22.9 p. 432) excretory canal Major Animal Phyla – Life cycle of human pork tapeworm (F22.10 p. 433) 22.3 What Are the Major Animal Phyla? • 22.3.4 Annelids Are Composed of Identical Segments – Figure 22.11 An annelid, the earthworm (p. 434) – Figure 22.12 Diverse annelids (p. 435) Copyright © 2005 Pearson Prentice Hall, Inc. nephridia coelom intestine excretory pore ventral nerve cord anus coelom hearts esophagus brain mouth ventral pharynx vessel ventral nerve cord crop gizzard intestine 22.3 What Are the Major Animal Phyla? • 22.3.5 Most Mollusks Have Shells – Figure 22.13 A generalized mollusk 435) – 22.3.5.1 Gastropods Are One-Footed Crawlers • Figure 22.14 The diversity of gastropod mollusks (p. 436) Copyright © 2005 Pearson Prentice Hall, Inc. (p. gonad heart digestive tract coelom tentacle shell ganglia (brain) mantle eye radula anus mouth gill foot nerve cords 22.3 What Are the Major Animal Phyla? – 22.3.5.2 Bivalves Are Filter Feeders • Figure 22.15 The diversity of bivalve mollusks (p. 436) Copyright © 2005 Pearson Prentice Hall, Inc. 22.3 What Are the Major Animal Phyla? – 22.3.5.3 Cephalopods Are Marine Predators • Figure 22.16 The diversity of cephalopod mollusks (p. 437) Copyright © 2005 Pearson Prentice Hall, Inc. 22.3 What Are the Major Animal Phyla? • 22.3.6 Arthropods Are the Dominant Animals on Earth – Figure 22.17 The exoskeleton allows precise movements (p. 438) – Figure 22.18 The exoskeleton must be molted periodically (p. 438) – Figure 22.19 Segments are fused and specialized in insects (p. 438) – Figure 22.20 Arthropods possess compound eyes (p. 439) Copyright © 2005 Pearson Prentice Hall, Inc. antennae head abdomen thorax compound eye mouth parts wing 22.3 What Are the Major Animal Phyla? – 22.3.6.1 Insects Are the Only Flying Invertebrates • Figure 22.21 The diversity of insects (p. 439) Copyright © 2005 Pearson Prentice Hall, Inc. 22.3 What Are the Major Animal Phyla? – 22.3.6.2 Most Arachnids Are Predatory Meat Eaters • Figure 22.22 The diversity of arachnids (p. 440) Copyright © 2005 Pearson Prentice Hall, Inc. 22.3 What Are the Major Animal Phyla? – 22.3.6.3 Myriapods Have Many Legs • Figure 22.23 The diversity of myriapods (p. 441) Copyright © 2005 Pearson Prentice Hall, Inc. (a) (b) 22.3 What Are the Major Animal Phyla? – 22.3.6.4 Most Crustaceans Are Aquatic • Figure 22.24 The diversity of crustaceans (p. 441) Copyright © 2005 Pearson Prentice Hall, Inc. 22.3 What Are the Major Animal Phyla? • 22.3.7 Roundworms Are Abundant and Mostly Tiny – Figure 22.25 A freshwater nematode (p. 442) – Figure 22.26 Some parasitic nematodes (p. 443) Copyright © 2005 Pearson Prentice Hall, Inc. anterior end mouth posterior end eggs vagina ovary cuticle intestine 22.3 What Are the Major Animal Phyla? • 22.3.8 Echinoderms Have a Calcium Carbonate Skeleton – Figure 22.27 The diversity of echinoderms (p. 443) – Figure 22.28 The water-vascular system of echinoderms (p. 444) Copyright © 2005 Pearson Prentice Hall, Inc. sieve plate ampulla tube feet canals plates of endoskeleton 22.3 What Are the Major Animal Phyla? • 22.3.9 The Chordates Include the Vertebrates Copyright © 2005 Pearson Prentice Hall, Inc.