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PHYLUM ECHINODERMATA (spiny skin) • 1. ECHINODERM CHARACTERISTICS They have pentaradial symmetry with the body divided into 5 (or multiples of 5) around the oral-aboral axis. –The advantage is they have sensory and feeding structures in all directions. 2. They have a skeleton made up of a series of plates called ossicles. The ossicles are held in place by connective tissue. 3. They have a water-vascular system consisting of water filled canals and the tube feet. It also contains a ring canal that surrounds the mouth. 4. The ring canal connects to the outside by a stone canal or a madreporite. The madreporite is used as an inlet to replace water. Polian vesicles are sac-like extensions of the ring canal that store fluid for use in the water-vascular system. 5. The radial canals branch off the ring canal and go out into the arms. The tube feet usually end in a bulb-like ampulla. When the ampulla contracts it squeezes water into the tube foot allowing it to move. 6. The tube feet have suckers which they use to attach themselves to the substrate. Some gas exchange and diffusion happens across the tube feet. 7. Class Asteroidea (seastars, star fish) Usually have five arms that radiate out from the central disk. 8. The mouth is located on the bottom of the disk, often surrounded by oral spines. Dermal branchiae are extensions of the body wall which extend out through the ossicles. They function in gas exchange (respiration). 9. A row of ossicles running the length of each arm on the oral surface (underside) is called the ambulacral groove. The groove consists of two tube feet with the radial canal and radial nerve in between. 10. Maintenance Functions. They feed on snails, bivalves, polycheates, coral, and detritus. 11. The stomach is very large and takes up most of the coelom. The stomach is divided into two parts. The cardiac stomach is the largest and receives the food first. 12. The aboral or pyloric stomach is used to distribute the nutrients to the rest of the body. 13. Most feed on bivalves. It does this by wrapping its arms around the bivalve and applying pressure. They use different arms as they tire until the bivalve weakens. 14. When the shells are about 0.1mm apart the starfish everts part of its cardiac stomach into the shell. It will secrete enzymes which begin to digest the bivalve inside its own shell. • When it weakens enough and can’t keep the shell closed the starfish finishes opening it and eats. 15. Gas exchange and removal of metabolic waste occurs by diffusion across the dermal branchiae, tube feet, and other membranes. The principle nitrogenous waste is ammonia. 16. The nervous system is composed of a nerve ring around the mouth and radial nerves that extend out to the arms and help control the tube feet. 17. They have sensory receptors located all over that help them respond to changes in light, temperature, chemicals, etc. Some have special tube feet on the ends of the arms that have ocelli (light sensitive cells). 18. Regeneration, Reproduction, and Development. If a starfish is damaged it is capable of regenerating the damaged parts. 19. Some perform asexual reproduction by dividing their central disks. • Most are dioecious. • External fertilization occurs when conditions are right. 20. The ciliated embryos float around eating plankton and become bipinnaria larva. When the arms start to grow it settles to the ocean floor and becomes a brachiolaria larva which develops into an adult starfish. 21. Sea Daisies Recently moved into class Asteroidea. They have no internal digestive system but a thin membrane called a velum is used to cover the substrate and digest and absorb all available nutrients. 22. CLASS OPHIUROIDEA (basket stars and brittle stars) a. Brittle stars have unbranched arms, basket stars have branched arms. 23. Most have no dermal branchiae and the tube feet have no suction cups. Tube feet are moved by muscles at their bases. Madreporites are located on the oral side. 24. They move about by using their arms to crawl over the substrate. 25. Maintenance Functions They are scavengers and predators feeding on plankton. Basket stars have mucus covering their tube feet which they wave around collecting plankton. 26. Regeneration, Reproduction, and Development. They have the capability to generate missing parts. In a process called autotomy they can use muscles to sever one of its own arms in an effort to escape. 27. They are dioecious. Fertilization is external. The larval stage is called an ophiopluteus. It swims and feeds on plankton. 28. CLASS ECHINODEA (sea urchins, sand dollars, heart urchins) SEA URCHIN Sea Urchin Sand Dollar Heart Urchin Heart Urchin 29. Sea urchins live on hard surfaces, sand dollars and heart urchins live on sand or mud. 30. Sea urchins are rounded, they have a skeleton called a test made up of ten plates that arch between the oral and aboral ends. Some are venomous. 31. Maintenance Functions They feed on algae, coral polyps, and scavenge dead animals. 32. The Aristotle’s lantern is a structure used for chewing. It’s function is to cut food into smaller pieces that can be eaten. They have a large coelom and nutrients are circulated in the coelomic fluids. 33. Reproduction, and Development – They are dioecious. – External fertilization. –An echinopluteus larva swims around for several months while it undergoes metamorphosis into an adult. 34. CLASS HOLOTHUROIDEA (sea cucumbers) – Elongated, no arms. – Tube feet surround the mouth and act like tentacles to gather food into the mouth. Sea Cucumber Sea Cucumber 35. 10 to 30 cm in length. • They have thick body walls with layers of longitudinal and circulatory muscles. They are eaten in some countries. • They use these muscles to wiggle along the sea floor. 36. Maintenance Functions. They eat by capturing food with their mucus covered tentacles. 37. They have a pair of tubes running the length of the body called respiratory trees. Their function is to remove nitrogenous wastes. 38. For defense their body can secrete toxins. They may also evert their respiratory trees then secrete a sticky mucus that entangles the predator and allows them to escape. 39. Reproduction and Development. • Dioecious • Fertilization is external. • They can also reproduce by transverse fission and regenerate lost parts. • 40. CLASS CRINOIDEA (sea lilies, feather stars) • The sea lily looks like a flowering plant but is an animal that also belongs to the echinoderm phylum. • 41. The sea lily has five or more arms but no spines. It does have hard plates in its skin and many tube feet. • Sea lilies live mostly on ocean floors. They use their round center stalks to attach themselves to the ocean floor. • 42. The mouths of the sea lily are on top of their bodies which are on top of the stalks. • Sea lilies have lived in the oceans for hundreds of millions of years making them the most primitive echinoderms • 43. the feather star is a close relative of the sea lily. • Unlike a sea lily, a feather star has no stalk. • It attaches itself to the ocean floor with many short hooks. • 44. A feather star spends much of its time attached to the ocean floor but it can also use its long arms to swim or to walk. • 45. both feather stars and sea lilies have tube feet that cover their long arms. The tube feet are coated with a sticky mucus. The mucus helps them capture food. • 46. feather stars and sea lilies eat plankton. When plankton gets caught on the tube feet it gets stuck in the mucus. The feather star then moves the plankton to a groove in the middle of its arm. • 47. from there the groove carries the food straight to the mouth of the echinoderm. Feather Star Sea Lilly • 48. Threats to Echinoderms • Starfish are keystone species in their respective marine communities. Their relatively large sizes, diverse diets and ability to adapt to different environments makes them ecologically important • The term "keystone species" was in fact first used by Robert Paine in 1966 to describe a starfish, Pisaster ochraceus. • The feeding activity of the omnivorous starfish Oreaster reticulatus on sandy and seagrass bottoms in the Virgin Islands appears to regulate the diversity, distribution and abundance of microorganisms. • These starfish engulf piles of sediment removing the surface films and algae adhering to the particles. • Organisms that dislike this disturbance are replaced by others better able to rapidly recolonise "clean" sediment. • In addition, foraging by these migratory starfish creates diverse patches of organic matter, which may play a role in the distribution and abundance of organisms such as fish, crabs and sea urchins that feed on the sediment. • Starfish sometimes have negative effects on ecosystems. Outbreaks of crown-of-thorns starfish have caused damage to coral reefs in Northeast Australia and French Polynesia • Several species may suffer a wasting disease caused by bacteria in the genus Vibrio. • Starfish and other echinoderms are sensitive to marine pollution. The common starfish is considered to be a bioindicator for marine ecosystems.