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Chapter 29 Section 1 Mollusks Phylum Mollusca • Snails, slugs, oysters, clams, scallops, octopuses, and squids are all mollusks. • Mollusks and annelids share a feature during the larval stage called a trochophore which develops from the fertilized egg. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks • In some species, the trochophore is freeswimming and propels itself through the water by movement of cilia on its surface. • The presence of a trochophore larva in mollusks and annelids suggests that they share a common ancestor. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Key Characteristics of Mollusks 1. The body cavity in mollusks is a true coelom, although in most species it is reduced to a small area immediately surrounding the heart. 2. Most mollusks exhibit bilateral symmetry. 3. Mollusks have organ systems for excretion, circulation, respiration, digestion, and reproduction. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Key Characteristics of Mollusks 4. The body of every mollusk has three distinct parts: the visceral mass, the mantle, and the muscular foot. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Key Characteristics of Mollusks • The visceral mass is a central section that contains the mollusk’s organs. • The mantle is a heavy fold of tissue that forms the outer layer of the body. • Finally, every mollusk has a muscular region called a foot, which is used primarily for locomotion. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Key Characteristics of Mollusks 5. Many mollusks have either one or two shells that serve as an exoskeleton, protecting their soft body. 6. All mollusks except bivalves have a radula, a tongue-like organ located in their mouth. The radula has thousands of pointed, backward-curving teeth arranged in rows. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Key Characteristics of Mollusks Organ Systems: Excretion • A mollusk’s coelom is a collecting place for wasteladen body fluids. • The beating of cilia pulls the fluid from the coelom into tiny tubular structures called nephridia. • The nephridia recover useful molecules (sugars, salts, and water) from the coelomic fluid. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Key Characteristics of Mollusks Organ Systems: Circulation • In a circulatory system, blood carries nutrients and oxygen to tissues and removes waste and carbon dioxide. • Most mollusks have a three-chambered heart and an open circulatory system. • Octopuses and squids are exceptions because they each have a closed circulatory system. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Key Characteristics of Mollusks Organ Systems: Respiration • Most mollusks respire with gills, which are located in the mantle cavity. • Most terrestrial snails have no gills. Instead, the thin membrane that lines their empty mantle cavity functions like a primitive lung. • Sea snails also lack gills, and gas exchange takes place directly through their skin. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Key Characteristics of Mollusks Organ Systems: Reproduction • Most species of mollusks have distinct male and female individuals, although some snails and slugs are hermaphrodites. • Certain species of oysters and sea slugs are able to change from one sex to the other and back again. • Many marine mollusks are moved from place to place as their trochophore larvae drift in the ocean currents. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Body Plans of Mollusks Gastropods- Stomach footed • Gastropods—snails and slugs—are primarily a marine group that has successfully invaded freshwater and terrestrial habitats. • Most gastropods have a pair of tentacles on their head with eyes often located at the tips. • Gastropods display varied feeding habits. Many are herbivores that scrape algae off rocks using their radula. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Characteristics of Gastropods Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Body Plans of Mollusks Bivalves • All bivalves have a two-part hinged shell. The valves, or shells, of a bivalve are secreted by the mantle. • Two thick muscles, the adductor muscles, connect the valves. When these muscles are contracted, they cause the valves to close tightly. • Bivalves are unique among the mollusks because they do not have a distinct head region or a radula. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Body Plans of Mollusks Bivalves • Many bivalves use their muscular foot to dig down into the sand. • Sexual Reproduction occurs by gametes being released into open water. (external fertilization) Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Characteristics of Bivalves Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Body Plans of Mollusks Cephalopods - Head footed • Squids, octopuses, cuttlefish, and nautiluses (only one to have an external shell) are all cephalopods. Most of their body is made up of a large head attached to tentacles. • Cephalopods are the most intelligent of all invertebrates. They have a complex nervous system that includes a well-developed brain. • The structure of a cephalopod eye is similar in many ways to that of a vertebrate eye, and some species have color vision. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 1 Mollusks Characteristics of Cephalopods •Most have a small internal shell or no shell at all •The Nautilus has only one external shell. •Move by jet propulsion •Most have a darkcolored foul tasting ink they can release when frightened. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 2 Annelids The First Segmented Animals • Annelids are easily recognized by their segments, which are visible as a series of ringlike structures along the length of their body. • Some of the segments are modified for specific functions, such as reproduction, feeding, or sensation. A well-developed cerebral ganglion, or primitive brain, is located in one anterior segment. • Internal body walls, called septa, separate the segments of most annelids. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 2 Annelids The First Segmented Animals Characteristics of Annelids 1. The fluid-filled coelom is large and is located entirely within the mesoderm. 2. The organ systems of annelids show a high degree of specialization and include a closed circulatory system and excretory structures called nephridia. 3. Most annelids have external bristles called setae. Some annelids also have fleshy appendages called parapodia. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 2 Annelids Annelid Groups Marine Worms • Marine segmented worms are members of class Polychaeta, the largest group of annelids. Polychaetes live in virtually all ocean habitats. • A distinctive characteristic of polychaetes is the pair of fleshy, paddle-like parapodia that occur on most of their segments. • The parapodia, which usually have setae, are used to swim, burrow, or crawl. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 2 Annelids Annelid Groups Marine Worms • Nereis, a polychaete worm, grasps its prey in its jaws, which open when it thrusts out its pharynx. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 2 Annelids Annelid Groups Earthworms • Earthworms and some related freshwater worms are members of the class Oligochaeta. Oligochaetes have no parapodia and only a few setae on each segment. • Earthworms lack the distinctive head region and have no eyes. • Their clitellum is involved in sexual reproduction. • Earthworms are highly specialized scavengers. They literally eat their way through the soil, consuming their own weight in soil every day. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Benefits of Earthworms • Their castings contain nutrients which are returned to the soil. • They aerate the soil as they move and burrow through it. • They break up the soil in which they live allowing plant roots to penetrate the soil. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 2 Annelids Anatomy of an Earthworm Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 2 Annelids Annelid Groups Hydrostatic Skeleton • The fluid within the coelom of each body segment creates a hydrostatic skeleton that supports the segment. • Each segment contains muscles that pull against this hydrostatic skeleton. • Circular muscles wrap around the segment, while longitudinal muscles span its length. Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Chapter 29 Section 2 Annelids Annelid Groups Class Hirudinea Leeches • Leeches are the only members of class Hirudinea. Leeches lack both setae and parapodia. • Most live in moist tropical countries • Usually no more than 6cm long and are somewhat flattened. • Most are freshwater that exist as external parasites few are marine or terrestrial • ¼ are carnivores not parasites and feed on soft-bodied invertebrates such as snails, worms, and insect larva • Drink blood and body fluids of their host Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Hirudinea characteristics • The body of a leech is flattened, and unlike other annelids, its segments are not separated internally. • A leech has suckers at both ends of its body. • Suckers at the anterior end attach the leech to the host. Suckers at the posterior end attach the leech to an object as it waits for a host to come by Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Leech Attachment • Leeches penetrate the skin of their host in one of two ways • Some have a muscular proboscis, a tubular organ that they force into the tissue of the host. • Other leeches slice the tissue with razor-sharp jaws. The use a muscular pharynx to suck blood out of the area. • Some can anesthetize the wound so the host does not know they have been bitten • Leeches release an anti-clotting agent to prevent the hosts blood from clotting Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved. Feeding • A meal can be as much as 10 times the weight of the leech • It can take the leech up to 200 days to digest its food. • A leech may need to feed only once a year Chapter menu Resources Copyright © by Holt, Rinehart and Winston. All rights reserved.