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WHAT’S GOING ON HERE??? Multiple Alleles Codominance Incomplete Dominance Sex-Linked Trait COMPLEX PATTERNS OF INHERITANCE When heredity breaks the rules… INCOMPLETE DOMINANCE Neither trait is dominant; the heterozygous individual is a BLEND of the two traits Example: In a variety of flowers (known as four o’clock flowers), crossing a homozygous red flower with a homozygous white flower makes heterozygous offspring that are PINK Problem: R = red R1 = white R R RR1 Parents: RR x R1R1 R1 RR1 Genotype: 4RR1 Phenotype: 4 Pink R1 RR1 RR1 CODOMINANCE BOTH alleles are expressed equally at the same time Example: Crossing a homozygous blackfeathered chicken with a homozygous whitefeathered chicken produces a black-and-white speckled chicken Problem: B = Black feathers W = White feathers Parents: BB x WW Genotype: 4 BW B B BW BW Phenotype: 4 black-andW BW white speckled W BW MULTIPLE ALLELES A trait is controlled by more than two alleles; results in multiple phenotypes Example: Blood types in humans – 4 possible phenotypes – Type A, B, AB, O Type A and Type B are CODOMINANT; Type O is RECESSIVE to A and B Problem: Cross a heterozygous Type B mom and a heterozygous Type A dad Parents: IBi x IAi Phenotype Possible Genotype Type A IAIA or IAi Type B IBIB or IBi IA IAIB IAi Type AB IAIB i IBi ii Type O ii IB i Genotype: 1 IAIB, 1 IAi, 1 IBi, 1 ii Phenotype: 1 Type AB, 1 Type A, 1 Type B, 1 Type O SEX LINKED TRAITS A trait controlled by a gene found ONLY on the XCHROMOSOME Each human has 23 pairs of chromosomes Pairs 1-22 are the same in males and females, and are called autosomes Pair 23 are the sex chromosomes which determine gender Females have two homologous X-chromosomes - XX Males have one X-chromosome (inherited from mom) and one Y-chromosome (inherited from dad) – XY Example: the gene for eye color in fruit flies (studied by Thomas H. Morgan) is found on the X-chromosome; RED is dominant and WHITE is recessive Problem: Cross a red-eyed male with a white-eyed female R = red r = white Parents: XRY x XrXr Xr Xr Genotype: 2 XRXr, 2XrY XR XRXr XRXr Phenotype: 2 red-eyed females r XrY 2 white-eyed males Y XY