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91 Figure 53—Percentage of each 4th-code hydrologic unit in the basin assessment area that includes centers of concentration of biodiversity, or of species rarity and endemism, of plants and animals (source: Marcot and others 1997). ICBEMP Centers of biodiversity, rarity and endemism are outside of biodiversity and endemism hot spots NOTE: Basin boundary 4th field hydrologic units State boundaries Centers of biodiversity, rarity and endemism Species > 60 − 80% Species > 80 − 100% Species > 40 − 60% Species 1 − 20% Species > 20 − 40% Species = 0% Legend Unique species assemblage components—The following integrity components relate to identifying unique vertebrate species assemblages by geographic area (ERU), as identified and listed in Marcot and others (1997).15 Many of these assemblages are of species that occur peripherally in the basin assessment area. Managing for the periphery of ranges of species can be important for maintaining long-term evolutionary potential (Furlow and Armijo-Prewitt 1995, Kruckeberg and Rabinowitz 1985, Lesica and Allendorf 1995), for it is at the edges of ranges where environmental factors often differ the most and new genotypes and phenotypes emerge (see Marcot and others 1997 for examples in the basin assessment area). Component (3a): Upper Klamath Basin species— We identified a set of 10 species that are closely associated with the Upper Klamath Basin ERU in southern Oregon (fig. 54). The database and map show number of species by 4th code HUC. Component (3b): Owyhee Upland species—We identified a set of nine species that are closely associated with the Owyhee Upland ERU in southern Oregon and Idaho (fig. 55). The database and map show number of species by 4th code HUC. Component (3c): Northern Great Basin species— We identified a set of three species that are closely associated with the northern Great Basin ERU in southern Oregon (fig. 56). The database and map show number of species by 4th code HUC. Component (3d): Amphibians of the Columbia River Gorge—This unique assemblage consists of amphibian species that enter into the basin assessment area mainly or solely through the Columbia River Gorge (fig. 57). The database and map show number of species by 4th code HUC. Component (3e): Snake River Headwaters ERU and western Wyoming species—A set of eight species were identified that are closely associated with the Snake River Headwaters ERU in southeastern Idaho, and the western Wyoming (Greater Yellowstone Ecosystem) region (fig. 58). The database and map show number of species by 4th code HUC. Component (3f): Upper Snake River ERU species—Two species were identified that are closely associated with the Upper Snake River ERU in southern Idaho (fig. 59). The database and map show number of species by 4th code HUC. Component (3g): Western Montana species— Two species were identified that are closely associated with western Montana (fig. 60). The database and map show number of species by 4th code HUC. Full trophic ladder component—Component (3h): Vertebrate carnivores—Eight wide-ranging vertebrate carnivores occur in the basin assessment area: grizzly bear (Ursus arctos), black bear (U. americanus), gray wolf (Canis lupus), cougar (Felis concolor), lynx (Lynx lynx), wolverine, fisher (Martes pennanti), and American marten (M. americana). Four of these species were mapped and are included in this component (fig. 61).16 The full set of vertebrate carnivores—and by inference, the full trophic ladder—once likely ranged throughout the entire basin assessment area, but now is present only in the northern Rocky Mountains, including Glacier National Park, Yellowstone National Park and adjacent lands, and in other locations where grizzly bear and gray wolf recovery areas overlap, as in central Idaho and north-central Washington. The database and map for this component show number of species by 4th code HUC (fig. 61). 15 Since the current analysis was conducted, some range maps of species have been refined, ranges of additional vertebrate species of the basin assessment area have been mapped, and additional species have been added to these lists of unique species assemblages by ERU and geographic area. The lists presented in Marcot and others (1997) reflect these additions but were not included in time to update analyses presented here. We do not anticipate, however, that these additions would qualitatively change results presented here. 92 16 Since this analysis, ranges for the remaining four carnivores also were mapped. Their distributional ranges more than overlap those of the four initially mapped carnivore species. Thus, their inclusion in this analysis does not change the results of identifying centers of high-carnivore richness. Text continued on page 101 93 Figure 54—Numbers of species closely associated with the Upper Klamath Basin ecology reporting unit (ERU), by 4th-code hydrologic unit in the basin assessment area (see Marcot and others 1997 for list of species). ICBEMP Basin boundary 4th field hydrologic units State boundaries Species = 10 Species = 4 Species = 8 Species = 3 Species = 1 Species = 2 Species = 0 Legend