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IB 362 Lecture 3 – Evolutionary and Ecological Principles Charles Darwin About time of voyages About time of Origin of Species H. M. S. Beagle 1831-1836 A l f r e d R u s s e l “Wallace’s Line” Helped found the field of biogeography Why are the organisms we see the way they are (structure, behavior, chemistry, etc.) Why do the organisms that we find living in a place live there, and not in other places? Why are some organisms more abundant than others in a given habitat? What determines which different organisms live together in the same community and which do not? Which is more important for determining the structure of communities: predation? competition? symbiosis? Ecology – the study of interactions between organisms and their environment, and the effects of these interactions on the distribution and abundance or organisms. Can be studied at different hierarchical levels: Biosphere Ecosystem Community Population Individual Evolution – heritable changes in organisms over time . Can be due to: Random drift in small populations Natural selection Immigration and emigration Mutations Molecular drive (not so relevant for our interests here!) Kinds of species interactions Territoriality +/- or -/- Competition +/- or -/- Predation +/- Commensalism +/0 Mutualism +/+ Parasitism +/- A couple of examples of more modern ecological and evolutionary studies Connell and Paine – Pacific intertidal communities Vermeij – impact of predation on morphology Joseph Connell UC Santa Barbara Robert Paine University of Washington North Pacific intertidal communities Sea anemone Dungeness crab zonation Dog whelk “loco” snail Barnacles seaweeds Predation and defense Grazing (herbivory) Territoriality in sea anemones Food webs Removal experiments Geerat Vermeij, UC Davis The role of predation in shaping gastropod shell morphology