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Ch.16 The Origin of Species • What Is a Species? • How Do New Species Form? – allopatric and sympatric speciation • How Is Reproductive Isolation Between Species Maintained? • What Causes Extinction? Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. What Is a Species? • Biologists Need a Clear Definition of Species • Species Are Groups of Interbreeding Populations • Appearance Can Be Misleading – Members of a species may differ in appearance (F16.1 p. 303) Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. How Do New Species Form? • Models of allopatric and sympatric speciation (F16.2 p. 304) • Allopatric Speciation (F16.3 p. 305) – By Geographic Separation of a Population • Sympatric Speciation (F16.4 p. 305) – By Ecological Isolation of a Population Copyright © 2005 Pearson Prentice Hall, Inc. Allopatric Sympatric original population original population geographical isolation ecological isolation genetic divergence genetic divergence reproductive isolation reproductive isolation Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. How Do New Species Form? • Changes in Chromosome Number Can Lead to Sympatric Speciation – Speciation by polyploidy Copyright © 2005 Pearson Prentice Hall, Inc. (F16.5 p. 306) Case 1: Selffertilization meiosis Case 2: Breeds with normal diploid parent fertilization meiosis viable, diploid gametes Copyright © 2005 Pearson Prentice Hall, Inc. meiosis fails; no viable gametes How Do New Species Form? • Change Over Time Within a Species Can Cause Apparent “Speciation” in the Fossil Record – Interpreting an evolutionary tree (F16.6 p. 307) Copyright © 2005 Pearson Prentice Hall, Inc. time present past Copyright © 2005 Pearson Prentice Hall, Inc. change Copyright © 2005 Pearson Prentice Hall, Inc. in form How Do New Species Form? • Under Some Conditions, Many New Species May Arise – Adaptive radiation Copyright © 2005 Pearson Prentice Hall, Inc. (F16.7 p. 308) Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. Maintaining Reproductive Isolation Between Species (T16.1,p. 310) • Premating Isolating Mechanisms Prevent Mating Between Species – Members of Different Species May Be Prevented from Meeting – Different Species May Occupy Different Habitats • Ecological isolation (F16.8 p. 309) – Different Species May Breed at Different Times • Temporal isolation (F16.9 p. 309) – Different Species May Have Different Courtship Rituals – Species’ Differing Sexual Organs May Foil Mating Attempts Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. Maintaining Reproductive Isolation Between Species (T16.1,p. 310) • Postmating Isolating Mechanisms Limit Hybrid Offspring – One Species’ Sperm May Fail To Fertilize Another Species’ Eggs – Hybrid Offspring May Survive Poorly – Hybrid Offspring May Be Infertile • Mechanisms of Reproductive Isolation Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. What Causes Extinction? • Localized Distribution & Overspecialization Make Species Vulnerable in Changing Environments – Very localized distribution can endanger a species (F16.10 p. 311) – Extreme specialization places species at risk (F16.11 p. 311) – Ethiopian wolves (FE16.1 p. 312) – Interactions with Other Organisms May Drive a Species to Extinction – Habitat Change and Destruction Are the Leading Causes of Extinction Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc. Copyright © 2005 Pearson Prentice Hall, Inc.