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Chapter 50 Introduction to Ecology and the Biosphere I. Study of Interactions Ecology and evolutionary biology Abiotic/Biotic factors Subfields – Organism -> population -> community -> ecosystem -> landscape -> biosphere Environmental Issues LE 50-2 Kangaroos/km2 > 20 10–20 5–10 1–5 0.1–1 < 0.1 Limits of distribution Tasmania LE 50-3 Organismal ecology Population ecology Community ecology Landscape ecology Ecosystem ecology II. Limits to Distribution Biogeographic realms – continental drift Dispersal and distribution Behavior and Habitat selection Biotic factors Abiotic factors Climate LE 50-5 Palearctic Nearctic Tropic of Cancer (23.5°N) Oriental Ethiopian Equator Neotropical (23.5°S) Tropic of Capricorn Australian LE 50-6 Species absent because Yes Area inaccessible or insufficient time Dispersal limits distribution? No Behavior limits distribution? Yes No Habitat selection Biotic factors (other species) limit distribution? Yes Predation, parasitism, Chemical competition, disease factors Water Oxygen Abiotic factors Salinity No limit pH distribution? Soil nutrients, etc. Temperature Physical Light factors Soil structure Fire Moisture, etc. LE 50-8 100 Sea urchin Seaweed cover (%) 80 Both limpets and urchins removed Only urchins removed 60 Limpet 40 Only limpets removed Control (both urchins and limpets present) 20 0 August 1982 February 1983 August 1983 February 1984 LE 50-10a North Pole 60°N Low angle of incoming sunlight 30°N Tropic of Cancer Sunlight directly overhead at equinoxes 0° (equator) Tropic of Capricorn 30°S Low angle of incoming sunlight 60°S South Pole Atmosphere LE 50-10b 60°N 30°N June solstice: Northern Hemisphere tilts toward sun; summer begins in Northern Hemisphere; winter begins in Southern Hemisphere. 0° (equator) March equinox: Equator faces sun directly; neither pole tilts toward sun; all regions on Earth experience 12 hours of daylight and 12 hours of darkness. 30°S Constant tilt of 23.5° September equinox: Equator faces sun directly; neither pole tilts toward sun; all regions on Earth experience 12 hours of daylight and 12 hours of darkness. December solstice: Northern Hemisphere tilts away from sun; winter begins in Northern Hemisphere; summer begins in Southern Hemisphere. LE 50-10c 60°N 30°N Descending dry air absorbs moisture 0° (equator) 30°S Ascending moist air releases moisture Descending dry air absorbs moisture 0° 60°S Arid zone Tropics Arid zone LE 50-10d Arctic Circle 60°N Westerlies 30°N Northeast trades Doldrums Southeast trades 0° (equator) 30°S Westerlies 60°S Antarctic Circle III. Aquatic Biomes Freshwater vs. Marine Zones – Photic – Aphotic – Benthic Distribution factors – – – – Thermocline Light penitration Distance from shore Open water vs. Bottom Lake depth (m) 0 4 Spring Winter O2 (mg/L) 8 12 8 16 2° 4° 4° 4° 4°C 24 O2 concentration 0° 4° 4° Lake depth (m) LE 50-13 O2 (mg/L) 0 4 8 12 8 12 8 16 24 4° 4° 4° 4°C High (>8 mg/L) Medium (4–8 mg/L) O2 (mg/L) 0 4 8 12 8 4° 4° 22° 20° 18° 8° 6° 5° 4°C 4° 4° 4° 4°C 16 24 Autumn Thermocline Summer Lake depth (m) Lake depth (m) Low (<4 mg/L) O2 (mg/L) 0 8 16 24 4 IV. Terrestrial Biomes Climate and climograph Vertical stratification Ecotone LE 50-18 Desert Temperate grassland Tropical forest 30 Temperate broadleaf forest 15 Coniferous forest 0 Arctic and alpine tundra –15 100 200 300 Annual mean precipitation (cm) 400 LE 50-19 30°N Tropic of Cancer Equator Tropic of Capricorn 30°S Key Tropical forest Savanna Desert Chaparral Temperate grassland Temperate broadleaf forest Coniferous forest Tundra High mountains Polar ice