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Palynology of the Devonian Jauf Formation from the northern part of Saudi Arabia • • • • • Mansour Al-Ruwaili, Saudi Aramco, Saudi Arabia Pierre Breuer, University of Liège, Belgium Ken Higgs, University of Cork, Ireland Philippe Steemans, University of Liège, Belgium Charles Wellman, University of Sheffield, UK Location of the boreholes After Loboziak, 2000 1700 feet or 520 m Stratigraphic position of the boreholes Y1 Y3 Y4 X1 X2 After Loboziak, 2000 After Al-Hajri et al., 1999 Palynological assemblages Palynological assemblages Continental palynomorphs • Palynological assemblages are strongly dominated by miospores (abundant trilete/monolete spores, and less frequent cryptospores) Palynological assemblages Marine palynomorphs • Rare acritarchs • One level contains numerous chitinozoans • Some levels contain large palynomorphs related to algae, or prasinophyceae? • D3b (leiospheridia bloom) 140 µm in diameter Biostratigraphy 1. Verrucosisporites polygonalis 2. Dictyotriletes subgranifer 3 2 3. Camarozonotriletes sextantii 1 After Streel et al., 1987 1. Synorisporites papillensis 2. Knoxisporites riondae AB or FD = early to middle Emsian ? After Streel et al., 1987 1. Rhabdosporites minutus After Streel et al., 1987 1. Clivosispora verrucata 2. Dictyotriletes emsiensis late FD or earliest AP = late Emsian After Streel et al., 1987 2. Grandispora protea 1. Acinosporites apiculatus 2 1 After Streel et al., 1987 1. Clivosispora verrucata 1 AP-Pro = latest Emsian After Streel et al., 1987 4. Geminospora lemurata 3. Grandispora velata 4 3 2 1 2. Grandispora gabesensis After Streel et al., 1987 1. Grandispora protea Biostratigraphic results Givetian ? Eifelian ? Emsian ? Emsian ? Pragian Biostratigraphic results ? ? 7 - 8 meters thick ? ? D3b Biostratigraphic results ? ? D3b ? ? Endemic species Examples of new species Examples of new species Examples of new species ? Cymbosporites n. sp. Dictyotriletes n. sp. A possible phylogeny? A possible phylogeny? A possible phylogeny? A possible phylogeny? A possible phylogeny? A possible phylogeny? A possible phylogeny? FD-Min Classe 4 FD-Pre Min Classe 4 Y4 Classe 1 Classe 2 Classe 3 Classe 4 Classe 3 Classe 2 Classe 1 2 3 A possible phylogeny? 1 4 Min AB + FD pre Min AB and FD Monolete species Previous records of the oldest monolete species • Arkhangelskaya 1985 - Devonomonoletes microtuberculatus n. g. & n. sp Limbomonoletes lenis n. g. & n. sp - Russia - late Emsian. • Kemp 1972 - Reticuloidosporites antarcticus n. g. & n. sp. - Antarctica – early Emsian Palaeogeography Palaeogeographic distribution of miospores Saudi Arabia Palaeogeographic distribution of K. riondae Palaeogeographic distribution of C. dammamensis Palaeogeographic distribution of miospores Saudi Arabia Conclusions • The Jauf Formation is Emsian in age. • The Emsian / Eifelian boundary is probably within the lowest part of the Jubah Formation. • The D3b event in north-western Saudi Arabia is precisely placed in the west European Min interval Biozone (mid Emsian). • The assemblages contain monolete spores which are among the oldest known with those from Antarctica. • A possible phylogeny could be defined ranging from a new Cymbosporites species to a new Dictyotriletes species. • Numerous new species have been observed. Some may allow definition of a north-western Gondwanan paleophytogeographic province.