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Development of tongue the tongue's embryonic origin is derived from all pharyngeal arches contributing different components. as the tongue develops "inside" the floor of the oral cavity, it is not readily visible in the external views of the embryonic stages of development. myogenesis (the development of muscular tissues) of the masticatory muscles are delayed in comparison with other muscles and are not complete even at birth, whereas the development of tongue muscles proceeds faster and ends at around birth Development of tongue begins at about 4th week of (IUL) due to local proliferation of mesenchyme in the floor of the mouth one elevation (swelling)appear in the Medline which is called median tongue or tuberculum median tongue it arise from endodermal floor of the pharynx this followed by two elevations called lingual projections or (bulges, swelling) distal tongue one on either side of the median elevation appear on the endodermal aspect of the mandibular processes Lateral lingual elevation quickly enlarge merge and fuse with each other and form large masse mucous membrane of anterior two third of tongue formed from this mass. Along the ventral and lateral margins of this elevation asulcus forms and deepened to form the linguogingival grooves. Floor of the mouth the tongue is separated by down growth of ectoderm around its periphery ,later on. this down growth of the ectoderm around the periphery of tongue separates the tongue from the floor of the mouth.(in this way lingual sulcus is formed and tongue becomes mobile) Foramen cecum Foramen cecum is found just behind the tuberculum impare.is related to the development of the thyroid gland .posterior to it ,the hypo brachial eminence is found ,which has two part a. Cranial part which give rise to posterior of the tongue b. caudal part .give rise to the epiglottis Tongue muscles originate from the mesoderm. The tongue muscle develops before masticatory muscles and is completed by birth. Masticatory muscles :originate from loose masses of the mesoderm. These muscles develop late and are not complete even at birth. Masticatory muscles develop muscle fibers within the tongue. And these muscles go through the same developmental changes as other skeletal muscle. The epithelium of the tongue is at first made up of single layer of cells, later it becomes stratified and papillae become evident. The line of the sulcus terminalis is marked by eight to twelve circumvallate papillae that develop at 25 months of the intrauterine life. The mucosa of the dorsal surface of the body of the tongue develops papillae at eleven weeks of the intrauterine life. At birth, the root mucosa becomes pitted by deep crypts that develop into lingual tonsils, the completion of which is marked by lymphocytic infiltration. Taste buds arise by the inductive interaction between epithelial cells and nerve cells in the tongue. Development of anomalies of tongue Examples are: Median rhomboid glossitis: it results from persistence of tuberculum impair. It is characterized by a red, smooth zone as seen in the midline, slightly anterior to the foramen caecum. Bifid tongue it is seen when the two lateral processes of the tongue are not fused completely. It is very rare Macroglossia (enlarged tongue): it is due to an over-development of the tongue. Microglossia (small tongue): it is due to an under-development of the tongue. Ankyloglossia: due to fusion of the tongue with the floor of the mouth. Mostly it is partial and very rarely complete . Fissured tongue: in this there are numerous small fissures or grooves on the dorsal surface of the tongue.