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Population Dynamics Population Dynamics • Changes in population size and the factors that regulate populations over time. Population Density • The number of individuals per unit area volume. volume • Example: the number of oak trees per km2 in a forest. Estimation of Population Size 1. 2. 3. Individual counts (not always practical) Transects Mark-recapture method: method N= marked individual X total catch second time recaptured marked individuals Example: pond turtles 1992: 18 1993: 34 (12) 1994: 30 (18) Estimation of Population Size Example: pond turtles 1992: 18 1993: 34 (12) 1994: 30 (18) 1992-93: N = 18(34) = 51 turtles 12 _________________________________________ 1993-94: N = 34(30) = 56.7 turtles 18 _________________________________________ So: 51 + 56.7 = 107.7 = 53.9 turtle estimation 2 Dispersion • The distribution of individuals within geographical population boundaries. boundaries • Three examples of dispersion patterns: 1. Clumped 2. Uniform 3. Random Dispersion Patterns 1. Clumped a. individuals are aggregated in patches b. unequal distribution of resources in the environment. Example: Example trees around a lake or pond Dispersion Patterns • 2. Uniform a. individuals are evenly distributed b. interactions among individuals of a population Example: creosote bushes in the desert Dispersion Patterns • 3. Random a. unevenly distributed b. random dispersion is rare Example: Example clams in a mud flat Understanding Population Growth 1. Exponential Growth Model a. exponential growth: rate of expansion (growth) of a population under ideal conditions. b. simple equation: G=rN G: growth rate of population N: number of individuals in the pop r: intrinsic rate of increase (range between -1 & +1) Exponential Growth • Example: bacteria produces a J-shaped curve Number of individuals (N) Time Understanding Population Growth 2. Logistic Growth Model a. logistic growth: environmental factors that restrict the growth of a population (called: population limiting factors) factors (K-N) b. equation: G=rN K G: N: r: K: growth rate of population number of individuals in the pop intrinsic rate of increase carrying capacity K: Carrying Capacity • The maximum stable population size that a particular environment can support over a relatively long period of time (K). (K) Logistic Growth • Example: turtles in a pond K produces a S-shaped curve Number of individuals (N) Time Question • What if? • N = almost 0: exponential growth (K/K = 1, then G=rN) • N = almost K (carrying capacity): growth rate is approaching zero population growth (ZPG) Factors That Limit Population Growth 1. Density-Dependent Factors: a. Population-limiting factors whose effects depend on population density. density b. The greater the pop, the greater the effects. Examples: 1. 2. 3. 4. Limited food supply Plagues Predation War Factors That Limit Population Growth 2. Density-Independent Factors: a. Population-limiting factors (abiotic) whose occurrence is not affected by pop density. density b. the greater the pop, the greater the effects Examples: 1. 2. 3. 4. Earthquakes Fires Hurricanes Freeze in the fall Boom and Bust Species • A rapid increase (boom) in a population followed by a sharp decline (bust). (bust) • Examples: a. Daphnia in a pond number of individuals in pop. boom Time bust Boom and Bust Species • Sometimes species (carnivore) depend on other species (prey) for food. • Example: Snowshoe hare and lynx number of individuals in pop. Years Evolution Shapes Life Histories • Life History: Series of events from birth through reproduction to death. death • Two basic types of life history strategies: 1. Opportunistic life history (r-selected species) 2. Equilibrial life history (K-selected species) 1. Opportunistic life history (r-selected species) Characteristics: 1. maturing time: 2. life span: 3. mortality rate: 4. times female is reproductive: 5. age at first reproduction: 6. size of offspring: 7. parental care: short short often high usually once early small none 2. Equilibrial life history (K-selected species) Characteristics: 1. maturing time: long 2. life span: long 3. mortality rate: often low 4. times female is reproductive: often many 5. age at first reproduction: late 6. size of offspring: large 7. parental care: often extensive Life History Examples r-selected species: 1. Garden weeds 2. Insects 3. Desert annuals K-selected species: 1. Humans 2. Oak trees 3. Elephants Survivorship Curves • Type 1 survivorship curve: High survival rates until old age. Example: humans % of survivors % of maximum life span Survivorship Curves • Type 2 survivorship curve: Intermediate between the extremes. extremes Example: % of survivors squirrel % of maximum life span Survivorship Curves • Type 3 survivorship curve: High mortality rates as larvae but decreased mortality at later ages. Example: % of survivors oysters frogs insects % of maximum life span Human Population Growth • Age structure Proportions of individuals of a population in different age groups. groups A typical population has three main age groups (age structure). 1. Pre-reproductive 2. Reproductive 3. Post-reproductive Age Structure Post-reproductive Reproductive Pre-reproductive Human Population Growth Today • Human population as a whole is growing exponentially. exponentially • Has doubled (doubling-time) three times in the last three centuries (doubled the carrying capacity several times). • Is now >6 billion, might reach ~8 billion by 2020. 2020 Question: • What are the reasons for the increase in human population? population • Answer: 1. Improved health 2. Technology 3. Decreased death rates Question: • What is Zero Population Growth (ZPG)? • Answer: a. birth rate equals death rate. rate b. intrinsic growth (r) = 0 Question: • What are two ways the human population can reach ZPG? 1. Limit the # of offspring per couple a. Reduces family size b. Voluntary contraception c. Family planning 2. Delay reproduction (late 20’s instead of early 20’s)