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White-tailed Eagle © Andrew Guppy Authors Andrew Stevenson Scottish Natural Heritage (SNH), Great Glen House, Leachkin Road, Inverness, IV3 8NW. [email protected] Rhian Evans Royal Society for the Protection of Birds, Tayside and Fife office, Ground Floor, Robertson House, 1 Whitefriars Crescent, Perth, PH2 0PA. [email protected] Summary • The Species Action Framework (SAF) partfunded the initial translocation and release of young white-tailed eagles (sea eagles) in the east of Scotland, as part of the overall reintroduction to Scotland of this species which started in the 1970s. • Through a partnership of Scottish Natural Heritage (SNH), Royal Society for the Protection of Birds (RSPB) Scotland and Forestry Commission Scotland (FCS), the East Scotland Sea Eagle Project (ESSE) released 85 birds between 2007 and 2012. • The ESSE project aims to help recover the historical range in Scotland and to found an additional population away from the west coast. • Restoring a large predator such as white-tailed eagle is a long-term project and requires continuing management and monitoring to ensure success. Introduction 1998 a further 58 eaglets from Norway were released in Wester Ross under a joint SNH and RSPB project. This release, combined with the increasing number of wild-bred young entering the population, had allowed the west coast breeding population to reach 42 pairs by 2007 when the third phase of releases in the east of Scotland commenced. Why was this species on the Species Action Framework list? The white-tailed eagle was an obvious candidate for inclusion under the SAF as a species for conservation action, as there was already a reintroduction programme underway. At the time of the SAF starting, consideration was being given by the Sea Eagle Project Team (now White-tailed Eagle Project Team) to a third phase of releases away from the west coast in order to enhance recovery of historical range, and to establish a second population to ensure a successful reintroduction. This resulted in SAF funding supporting the ESSE project, under which translocated eaglets from Norway were released in Fife. Species background This section draws on the following key references: Love (1983, 2007). The white-tailed eagle (Haliaeetus albicilla), also known as the sea eagle, is a native species to Scotland. However, by the early 20th century it had become extinct through persecution, primarily owing to the belief that it was a major predator of livestock and game. Across Europe the species was also in decline and it became a very rare visitor to Scotland with little prospect of natural recolonisation. Two independent releases, each of four birds, at Loch Etive, Argyll in 1959 and on Fair Isle, Shetland in 1968, initiated thoughts of a more formal reintroduction. In the early 1970s the Nature Conservancy Council decided to undertake a sustained chick release programme. From 1975 to 1985, 82 eaglets from Norway were released on Rum National Nature Reserve. This release established territorial pairs on several west coast islands with the first successful breeding occurring on Mull in 1985. This initial population relied on a few consistently successful pairs and there were concerns about the long-term viability of the population. A second phase of releases was agreed to bolster the population. From 1993 to Habitat, distribution and abundance The species is widespread across much of northern Eurasia, breeding from Greenland east to Japan, and south into south-east Europe, the Middle East and central Asia (see white-tailed eagle factsheet for more information on population size and distribution). When the Scottish reintroduction started in the 1970s the species was listed as being Near-Threatened by IUCN, however conservation efforts across its range lead to its status being revised to Least Concern in 2006. As indicated by its large geographical range the species occupies a wide range of habitats. There is an association with coastal and freshwater habitats in both lowland and upland situations. It is also found in both open and wooded or forest habitats. In many areas where its abundance is relatively high it lives close to human populations including winter roosts and nesting pairs near large towns and cities. The species is also adapted to a wide range of climatic conditions, again evidenced by its broad distribution, though some northern populations are migratory. General ecology In Scotland comparison is often made with the golden eagle (Aquila chystaeos), however the white-tailed eagle has significant differences in its ecology and several studies in Scotland indicate that there is minimal competition between the two species (Evans et al., 2010; Whitfield et al., 2013). White-tailed eagles as adult pairs can be less territorial and can nest at fairly high densities more often than golden eagles. This more social behaviour extends to the immature birds too. As with other large raptors, young white-tailed eagles take around five to six years to reach breeding age. During this period they range widely but also form non-breeding communal roosts. These roosts may have an important role in pair formation amongst older immature birds in particular. The species has a relatively long breeding season, laying eggs during March and April, with young leaving the nest usually in July and August. Adult pairs generally remain on territory all year, roosting at or near the nest although they will at times join nearby communal roosts too. Nest sites in Scotland are usually in trees or on cliffs and crags. Some of the latter sites are almost ground nests whilst a range of tree species are used, though most are in conifers. The main requirements for tree nests are that the tree can support the large stick-nest and has limited exposure to adverse weather. The diet of the species is wide, including both live prey and carrion. They generally take a greater proportion of waterbird and seabird prey compared to golden eagles (Whitfield et al., 2013). Most prey consist of medium-sized mammals (e.g. rabbits and hares) and birds (e.g. gulls, ducks, geese, auks, grouse and, especially in Scottish coastal sites, fulmar). In some parts of their world range they eat mainly fish. In Scotland fish form a smaller component of the diet although over 20 species have been recorded in the diet of the reintroduced population (Marquiss et al., 2004). Most carrion consumed in Scotland is deer or sheep. History of decline, contributory factors and current threats The species became extinct in Scotland in 1918 following a decline which was largely attributed to persecution. The species was considered a serious predator of livestock especially sheep and was heavily persecuted in the 19th and early 20th centuries, with specimen and egg collectors also contributing to the eventual extinction as the species became rarer. Prior to the reintroduction starting in the 1970s, consideration was given to how viable it would be. The factors that contributed to the decline were considered before it went ahead and the detailed monitoring of the population since reintroduction has helped identify and manage threats and risks to the species. The main threats historically of persecution and egg collection though still present today have declined in importance. The first nesting pairs in the west were very intensively monitored with volunteer nest-watches to try to prevent interference with the birds. These were successful and fundamental in allowing the population to become established. The perception of white-tailed eagles as predators of lambs still prevails, and has been a difficult one to deal with at times. SNH has responded to concerns from the farming community and their representative bodies about alleged lamb predation throughout the reintroduction. Two studies for SNH into lamb predation, one on the island of Mull where white-tailed eagles first bred, and a later one at Gairloch in Wester Ross, identified that the eagles do take a small number of live lambs, but the majority of lambs taken were scavenged as carrion (Marquiss et al., 2004; Simms et al., 2010). As result of this research, a Sea Eagle Management Scheme has been established by SNH to help those farmers and crofters at risk of losing some lambs from eagle predation. Funding is available to both help improve lamb and sheep flock health and to undertake preventative measures so they are less likely to be impacted by eagles, rather than to compensate for losses. This has helped allay the concerns of some but not all. White-tailed eagles continue to be persecuted in Scotland, as a number of reintroduced eagles are known to have been killed illegally or to have disappeared in suspicious circumstances inferring illegal killing. Whilst these losses may have had some influence on the establishment and growth of the population, they have not been at a level that has hampered the re-establishment of the species. A newer potential threat is from wind farms, as information from Scandinavia and continental Europe has shown that white-tailed eagles are relatively poor at avoiding turbines and so are at risk of collision (May et al., 2010; Dürr, 2015). In Scotland, to date, risks to white-tailed eagles have been minimised through discouraging developments in key areas. Both persecution and wind farm collision risk are of most concern where they may affect new ‘pioneering’ pairs outside the main range, as loss of these pairs could impact on range recovery. Disturbance of nesting pairs and of regular roosts can also be an issue, and guidance has been drawn up by Forestry Commission Scotland to help minimise the effect of operations on white-tailed eagle roost and nest sites in commercial forests. This guidance can also be applied to a wide range of activities that occur outwith forestry (see Further Information). Over 200 eyries were monitored annually by NOF in three counties to allow enough donor chicks to be identified and collected. • A release site was secured and holding aviaries constructed. • Young eagles were translocated in June and released in August each year from 2007 to 2012. Eighty-five birds were released in total. Planning Aims Aims for 2007-2012 • To translocate and successfully release 15-20 white-tailed eagles per annum to the Tay/Forth area between 2007 and 2012. • To monitor released birds as they establish territorial and ultimately nesting pairs. The aim post-2012 has been: • To augment the range expansion of the west coast population of white-tailed eagles by establishing a viable, self-sustaining breeding population in the east of Scotland. Management Action Summary of the main action undertaken • A partnership was formed between RSPB Scotland, SNH and FCS, with RSPB Scotland and SNH jointly funding the project between 2007 and 2012. RSPB Scotland employed a project officer, and FCS provided the release site. Through this partnership, the East Scotland Sea Eagle (ESSE) project steering group was also formed. • Appropriate assessments were made and the necessary parties consulted. • Donor stock was identified in Norway and agreement made with Norwegian Birdlife partner Norsk Ornitologisk Forening (NOF). The decision to continue with the third phase of reintroduction to Scotland was made by the Sea Eagle Project Team in 2003. Research concluded that 85-100 white-tailed eagle chicks would be necessary to establish a new breeding population in eastern Scotland. A feasibility study and habitat assessments were carried out for several suitable geographical areas in Scotland for the reintroduction site prior to the final site selection. The project was fully considered against IUCN guidelines (IUCN, 1998). Once the final release site had been chosen RSPB Scotland staff, on behalf of the ESSE project steering group, also liaised with potential stakeholders and public bodies that may be affected by the reintroduction, including the bodies representing landowners, agricultural interests and elected representatives. In June 2007, the then Minister for Environment, Michael Russell MSP, approved the licence for the reintroduction programme. Location of work Although other areas were considered, the Tay/ Forth estuary area was concluded as being the most suitable location for the third phase of the reintroduction. It provided the largest estuary area and much high quality habitat for feeding and nesting. This area also had a relatively low incidence and risk of illegal persecution of birds of prey, and minimal potential conflict with sheep farming, compared to other sites considered. Additionally, though not part of the formal assessment, a high proportion of Scotland’s people live close by, providing opportunities to encounter the birds with substantial educational and eco-tourism potential. FCS provided a suitable secure aviary site at Weddersbie Forest, a quiet and relatively isolated woodland with minimal recreational or operational disturbance. The site was kept secret, though regular users knew and respected its sensitivities. This also gave the local community a sense of ownership and acted as a security measure in itself. The aviaries were monitored using CCTV cameras with images sent to a device monitored by the project officer. Local landowners were positive and cooperative, and a local farmer assisted with access and allowed project staff to store food for the captive eagles. Fife Police checked the site regularly. Methods and techniques A donor stock of young white-tailed eagles was identified through the Norwegian Sea Eagle Project. Licences were granted for chicks to be collected in three counties: Møre & Romsdal, Hordaland and Sogn & Fjordane. Licences required for the exportation, import, holding and release of the birds were necessary under CITES (for export and import), Animal Health legislation, and the Wildlife and Countryside Act 1981. A project officer was employed in April 2007 prior to the first cohort of white-tailed eagles being collected. Aviaries were designed and constructed with an artificial nest platform, perches, and a large release hatch in the front with thick, double layer mesh (Fig. 1). A door and feeding hatch were located in the back of the aviary allowing staff to feed the birds without unnecessary contact. Nest platforms were 1 m above the ground and lined with untreated tree bark and a layer of sphagnum moss. Staff from the Norwegian Sea Eagle Project monitored active sites in January to identify potential donor nests in June. Chicks were collected only from nests containing twins or triplets, at the age of 5-8 weeks, when they were old enough to regulate their own body temperature and feed themselves unassisted by parent birds, but small enough not to abandon the nest. Prior to export, the chicks were checked by vets to ensure that they were fit to be transported. This also satisfied the requirements of the CITES export and import licences, and the Animal Health import licence. In 2007, the Royal Norwegian Air Force transported the chicks to RAF Kinloss. In subsequent years, an aircraft was chartered to transport the chicks to Edinburgh airport. After the relevant customs procedures and paperwork had been processed airside, the chicks were transferred in vehicles to the aviaries. On arrival at the aviaries, health screening was carried out in conjunction with Natural Research Ltd (NR) and the Exotic Animal and Wildlife Service (EAWS) of the University of Edinburgh’s veterinary school. Full haematological and biochemical analyses were carried out, and samples taken to monitor lead levels, faecal parasites and faecal bacterial cultures. Blood smears were also taken for haemoparasite analyses and blood plasma was stored for further analyses. All birds were sexed by NR using DNA from blood samples. Chicks were weighed and biometric measurements taken. The chicks were then housed in pairs and matched as closely as possible according to size to prevent competition and exclusion for food. The chicks were fed twice a day initially, on a varied diet of fish, rabbit, venison and grey squirrel. All the food was donated by FCS, local game dealers, fish shops, and the Scottish Wildlife Trust who provided culled grey squirrels. The chicks were medicated to prevent the potential development of stress-related illnesses for the first five days in captivity, and vitamin and mineral supplements were added to their food throughout the captive period. Feeds were reduced to once a day as the birds grew and were able to tear apart larger prey. Fig 1. An example of an aviary used to house the young white-tailed eagles. © Andy Hay/RSPB Images The chicks were kept in captivity for 7–8 weeks and released when they were roughly 14 weeks old to coincide with when they would naturally fledge the nest. A week prior to release, all chicks were re-sampled for health parameters and given a welfare check by vets. Wing and tail-feather growth was checked and full biometrics taken again. Every chick was fitted with tags on each wing to enable identification and ageing of birds in the field (Fig. 2). Different colours were used each year with a single letter or number legend. Due to a change in the licensing procedure, wing tags were not fitted in 2008. The chicks were also fitted with 70 g Biotrack VHF transmitter backpacks using a Teflon harness authorised by BTO (Fig. 2). Each bird had a unique frequency so that dispersal could be monitored. The VHF transmitters have a mortality setting which is triggered if a bird is stationary for more than 6 hours, allowing the retrieval of any dead birds for post-mortem examination. Fig 3. A young white-tailed eagle being released in August 2010. © Andy Hay/RSPB Images During the winter months, the project officer gave talks to local interest groups and attended events to raise awareness of the project, and to engage with the community close to the release site and beyond. Fig 2. A young white-tailed eagle with a newly fitted VHF transmitter and wing tags. In 2010 a part-time education officer was employed to deliver a programme for primary schools in Tayside and Fife, to engage local children in the project and wider conservation. Resources were created for teachers and information was shared on the Scottish schools intranet ‘Glow’. Links with schools in Norway were also encouraged. Volunteer field teachers also played a key role in successfully delivering this programme. © Andy Hay/RSPB Images The releases were staggered over a period of a week of dry weather, and project partners, stakeholders, volunteers and media were invited to attend (Fig. 3). A food dump was maintained at the site for the first three months following release. All birds were monitored through systematic radiotracking and following up of sightings from the public. Every contact or sighting of a released bird was recorded in a database. Problems and solutions Since the first release in 2007, there have been 14 reported incidents of released white-tailed eagles predating on domestic poultry and wildfowl, and five cases of game bird predation, varying in severity. All cases involved juvenile birds, and occurred up to eight weeks after release. Two incidents of white-tailed eagles scavenging lamb carcasses occurred, but the lambs were already dead. Every incident was followed up immediately by the project officer and measures were taken to prevent further incident through positive management. This was done with advice and collaboration with SNH area officers. Each year during the project there were also several cases of mortality (see Fig. 4). A protocol for retrieving carcasses and appropriate post mortem or toxicology was agreed between EAWS and NR, RSPB Scotland, Tayside and Fife Wildlife Crime Officers, Science and Advice for Scottish Agriculture (SASA) and Scotland’s Rural College (SRUC). Mortality of released birds was caused by collisions with wires/electrocution, collisions with trains and persecution, and one bird was euthanized after being found with severe injuries from being struck by a seal on the Isle of May. In 2014, a three year old male was killed by a wind turbine in the Ochil Hills, Clackmannanshire. Dead birds were detected through radio-tracking or reported by members of the public. The majority of fatalities were juveniles and occurred between the release period and their first winter. Young birds were occasionally found unwell or injured and in need of rehabilitation, which was undertaken by Alistair Lawrie and the Scottish Society for the Prevention of Cruelty to Animals (SSPCA) before re-release. During the first two years of the project, some of the VHF transmitters failed to work. Monitoring of these individuals was therefore difficult, but still possible through observation of wing tags. Through the media, critics of the project questioned the viability of the reintroduction. Some doubted the suitability of the Scottish climate, claiming that it is too wet for white-tailed eagles. However, the survival rates of the released birds are similar to those of previous Scottish releases and some wild populations elsewhere (Evans et al., 2009). Others assumed that the birds were tame due to the belief that they had been imprinted during their time in captivity. However, considerable care was taken to avoid human contact before birds were released and this species appears more naturally ‘tolerant’ of human activities than other large raptors. Partnership working and resourcing The ESSE project was managed via regular Steering Group meetings throughout the project. Regular reports were made to the Scottish Government. As well as supporting the project financially, SNH also assisted with licensing and assessed reported livestock incidents. Additional funding for the final year of releases in 2012 came from Heritage Lottery Fund (HLF) and Rural Tayside and Fife LEADER. Volunteers played a key role with activities such as feeding birds in captivity, stocking the food dump, post-release monitoring of the birds and data entry. One volunteer used her expertise to make a short documentary about the project for schools. Outwith the area covered by radio-tracking, sightings from the public provided invaluable information about the birds’ dispersal. Results Between 2007 and 2012, 85 birds were released into the wild in Fife. The overall survival rate of the released birds to the end of 2014 was 69% with 27 carcasses recovered. This was comparable to the mortality rate seen in the west coast-released population. Fig. 4 summarises the survival statistics for 2007–2014. Fig. 5 shows the dispersal of east coast-released white-tailed eagles and indictes where most birds have spent their time. Fig 4. Summary of survival and causes of mortality of white-tailed eagles released in the east of Scotland between 2007 and 2012, as of December 2014. White-tailed eagle release year 2007 2008 2009 2010 2011 Number of Individuals 1 eagle 2-7 eagles Fig 5. Approximate distribution of east coast-released white-tailed eagles after their release. © RSPB Lessons Learnt, Further Work and Future Recommendations Lessons learnt The east coast project has built on west coast experience of rearing and releasing. The design of the release aviaries was based on those used during the successful second phase of releases in Wester Ross, and previous red kite releases in Scotland. The east coast project also had the advantage of monitoring all birds using radio-telemetry. This gave information on dispersal and survival. Better technology, such as satellite tags, has provided more accurate information about bird movements and roost sites and allows rapid mortality detection, though at much greater cost. Due to the location of the release site, the east coast project had the benefit of more accessible expert veterinary care than the more remote west coast releases. Welfare techniques have also improved since the initial release on Rum and the methods that are now used minimise human contact as much as possible. As the first nesting attempts of the east coastreleased birds occur, there will be careful management of the threats outlined previously and of land management activities close to nests to ensure the success of their establishment in the east of Scotland. Constructive engagement with the relevant stakeholders will be essential to achieve this. The value of volunteers, stakeholder engagement and the help of the public in providing sightings of the birds has been immense. Without these, the project would not have been a success. The project was fortunate in securing a release site in FCS woodland with a cooperative neighbouring farmer and community. The farmer lent his support throughout. The support from the local community has been paramount to the security at the release site, with several members reporting sightings of the newly released birds. Overall, the east coast white-tailed eagle project highlighted the need for careful planning, and for strong representation from Scottish and international project partners to overcome management issues and potential criticism. Key Management Messages Five key elements of the project can be identified: • Meticulous planning and coordination throughout. • Strong partnership and overseas connections. • A well coordinated post-release monitoring scheme. • Sympathetic, reactive, and constructive approaches to management issues. • Applying knowledge from previous successful release methods and post-release monitoring Future recommendations • The methods and techniques for releasing birds used during this project have been successful, and therefore could be considered for similar projects elsewhere with the provision that local conditions are taken into account. • The consultation process should be thorough, and ensuing monitoring should meet strict IUCN guidelines on reintroductions, especially in a densely populated area where the risk to the reintroduced population may be greater. • Strong partnerships are essential. Planning and preparation should be thorough, and communication should be maintained with partners and wider stakeholders throughout. • A contingency should be put in place for scenarios such as wildlife crime incidents, livestock predation, and conflict with land management activities. Positive management solutions should be offered as a proactive measure as well as preparing for a reactive and sympathetic response. New and ongoing work since SAF ended • Since the SAF project funding came to an end in 2011, the partnership between RSPB Scotland, SNH and FCS has remained. Funding was secured from HLF and Leader for a further year of translocation, allowing six more birds to be released. • Steering group meetings continued beyond the translocation phase of the project to address further management issues involving the released birds and establishing pairs. A volunteer network has been formed to help monitor these new pairs and to help protect nesting attempts. • Monitoring of the released birds continued in accordance with IUCN guidelines through a programme of radio-tracking. A database to record all contact with the released birds, and for future analysis of dispersal, is still maintained. • In 2013 the first pair of released birds from the east coast project bred successfully in Fife, with four pairs established in the east by 2014. Other east coast pairs have established and some individuals have successfully bred with wild-fledged west coast birds. The overall Scottish population had increased to 98 pairs in 2014. Further Information • http://www.birdlife.org/datazone/ speciesfactsheet.php?id=3364 – information on global population size and distribution of whitetailed eagle, from Birdlife International. • http://www.rspb.org.uk/ discoverandenjoynature/discoverandlearn/ sea-eagles/index.aspx – information about the East Coast project on the RSPB website. • http://www.rspb.org.uk/community/ourwork/b/ eastscotlandeagles/default.aspx – the RSPB’s ESSE project officer’s blog. • http://www.snh.gov.uk/about-scotlands-nature/ species/recent-species-projects/sea-eagle/ – information about white-tailed eagle on the SNH website. • http://www.snh.gov.uk/land-and-sea/ managing-wildlife/sea-eagle-managementscheme/ – information on the SNH Sea Eagle Management Scheme. • http://www.iucn.org/news_homepage/ news_by_date/?13377/New-Guidelines-onconservation-translocations-published-by-IUCN – information on the revised IUCN Guidelines published in 2013. • http://www.forestry.gov.uk/pdf/fcpn101. pdf/$file/fcpn101.pdf – FCS Practice Note on Managing forests for white-tailed eagles. • http://www.snh.gov.uk/protecting-scotlandsnature/reintroducing-native-species/scct/ – information on the new Scottish Code for Conservation Translocations, published in 2014. • Love JA. 2013. A Saga of Sea Eagles – a revised edition of Love’s 1983 book with further detailed accounts of phase 2, east coast and Irish reintroductions. Whittles Publishing: Dunbeath. Love JA. 1983. The return of the Sea Eagle. Cambridge University Press: Cambridge. Love JA. 2007. White-tailed eagle. In: The Birds of Scotland, Forrester RW, Andrews IJ, McInerny CJ, Murray RD, McGowan RY, Zonfrillo B, Betts MW, Jardine DJ, Grundy DS (eds). Scottish Ornithologists’ Club: Aberlady; 451-455. Marquiss M, Madders M, Irvine J, Carss DN. 2004 The Impact of White-tailed Eagles on Sheep Farming on Mull. Final Report. Centre for Ecology and Hydrology, Banchory. May R, Hoel PL, Langston R, Dahl EL, Bevanger K, Reitan O, Nygård T, Pedersen HC, Røskaft E, Stokke BG. 2010. Collision risk in white-tailed eagles: Modelling collision risk using vantage point observations in Smøla wind-power plant. NINA Report 639. Norwegian Institute for Nature Research, Trondheim. Simms IC, Ormston CM, Somerwill KE, Cairns CL, Tobin FR, Judge J, Tomlinson A. 2010. A pilot study into sea eagle predation on lambs in the Gairloch area – final report. Scottish Natural Heritage Commissioned Report 370, Battleby, Perth. Whitfield DP, Marquiss M, Reid R, Grant J, Tingay R, Evans RJ. 2013. Breeding season diets of sympatric White-tailed Eagles and Golden Eagles in Scotland: no evidence for competitive effects. Bird Study 60: 67-76. References Dürr T. 2015. European Bird Collision reporting spreadsheet: bird fatalities at wind turbines in Europe, at 1 June 2015 (available at http:// www.lugv.brandenburg.de/cms/detail.php/ bb1.c.312579.de. Accessed 28 July 2015. Evans RJ, Wilson JD, Amar A, Douse A, Maclennan A, Ratcliffe N, Whitfield DP. 2009. Growth and demography of a re-introduced population of White-tailed Eagles Haliaeetus albicilla. Ibis 151: 244–254. Evans RJ, Pearce-Higgins J, Whitfield DP, Grant JR, MacLennan A, Reid R. 2010. Comparative nest habitat characteristics of sympatric White-tailed Haliaeetus albicilla and Golden Eagles Aquila chrysaetos in western Scotland. Bird Study 57: 437-482. IUCN. 1998. Guidelines for Reintroductions. IUCN/ SSC Reintroduction Specialist Group, IUCN, Gland, Switzerland and Cambridge. Acknowledgements We thank Alv Ottar Folkestad and members of the NOF, RSPB, FCS and SNH staff, the Sea Eagle Project Team, and acknowledge the support and advice of west coast RSPB staff, the Scottish Government, the Norwegian Government, Jo Hedley, Alistair Lawrie, staff at the Lawrie Veterinary Group and the SSPCA’s national wildlife rehabilitation centre for care of sick and injured birds. We also thank Natural Research, Tayside and Fife Wildlife Crime Officers, Anna Meredith and Joanna Hedley at EAWS, Air Partner and Air Charters, our release site neighbours, particularly Neil and Lesley Powrie, and all the volunteers who devoted so much time and hard work to the project. Local businesses in Tayside who provided food for the captive eagles and the food dump. We are also very grateful to the Heritage Lottery Fund, Rural Tayside and Fife Leader for additional funding in 2012. Phil Whitfield provided invaluable comments on the draft. The SAF Partners • Scottish Natural Heritage • RSPB Scotland • Forestry Commission Scotland The Species Action Framework Handbook This account comes from the Species Action Framework Handbook published by Scottish Natural Heritage. For more information on the handbook please go to www.snh.gov.uk/ speciesactionframework. This document should be cited as follows: Stevenson A, Evans, R. 2015. White-tailed eagle. Version 1.0. In The Species Action Framework Handbook, Gaywood MJ, Boon PJ, Thompson DBA, Strachan IM (eds). Scottish Natural Heritage, Battleby, Perth.