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Interspecific Competition
• Superior and inferior
competitor
• Rates of reproduction
• Rates of growth
• Tolerance to limiting factors
• Direct competition
Mechanisms for superior competition
•
•
•
•
Consumption
Preemption
Overgrowth
Chemical interactions
(allelopathy)
• Territoriality
• Encounter competition
Competitive Exclusion
Eastern cottontail
New England cottontail
Coexistence of Groton rodents
How can six species of salamander coexist on
Groton School lands?
Coexistence
Resource Partitioning
Resource-based models
Spatially based models
Temporal Models
Changes over time (long-term)
Low levels of competition allow for extended
persistence
Limiting Factor Models
Predation
Disease
Parasitism
Competition and Niche
• Fundamental Niche
• Realized Niche
• Resource Partitioning
– Spatial
– Resource Selection
– Timing?
Spatial heterogeneity and variation
•
•
•
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Environmental Gradients
Disturbance
Soil/bedrock
Aspect/topography
Temporal Variation
• Parameters that vary
predictably over time
– Temperature
– Moisture
– Seed and nut production
– Prey population cycles
• Response of the inferior
competitor
– Dispersal
– Dormancy
– Reduction in density
Predation and Herbivory
Competition and Evolution
• Short-term changes
– Behavioral
• Long-term changes
– Structural
• Character
Displacement
• Behavioral plasticity
How do so many New England carnivores
persist in a small area?
Full Circle
• Why are the invasive species so successful in this system?
• How do so many New England carnivores persist in a small area?
• How can six species of salamander coexist on Groton School
lands?
Competition and Conservation
Small Whorled Pogonia
Isotria medeoloides
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