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Respiration Chapter 39 Part 1 Impacts, Issues Up in Smoke Smoking immobilizes ciliated cells and kills white blood cells that defend the respiratory system; highly addictive nicotine discourages quitting 39.1 The Nature of Respiration All animals must supply their cells with oxygen and rid their body of carbon dioxide Respiration • The physiological process by which an animal exchanges oxygen and carbon dioxide with its environment Interactions with Other Organ Systems Fig. 39-2b, p. 682 food, water intake oxygen intake Digestive System Respiratory System nutrients, water, salts oxygen elimination of carbon dioxide carbon dioxide Circulatory System Urinary System water, solutes elimination of food residues rapid transport to and from all living cells elimination of excess water, salts, wastes Fig. 39-2b, p. 682 Partial Pressure Partial pressure • Of the total atmospheric pressure measured by a mercury barometer (760 mm Hg), O2 contributes 21% (160 mm Hg) 760 mm Hg Fig. 39-3, p. 682 The Basis of Gas Exchange Respiration depends on diffusion of gaseous oxygen (O2) and carbon dioxide (CO2) down their concentration gradients Gases enter and leave the internal environment across a thin, moist layer (respiratory surface) that dissolves the gases Factors Affecting Diffusion Rates Factors that increase diffusion of gases across a respiratory surface: • High partial pressure gradient of a gas across the respiratory surface • High surface-to-volume ratio • High ventilation rate (movement of air or water across the respiratory surface) Respiratory Proteins Respiratory proteins contain one or more metal ions that reversibly bind to oxygen atoms • Hemoglobin: An iron-containing respiratory protein found in vertebrate red blood cells • Myoglobin: A respiratory protein found in muscles of vertebrates and some invertebrates 39.2 Gasping for Oxygen Rising water temperatures, slowing streams, and organic pollutants reduce the dissolved oxygen (DO) available for aquatic species 39.1-39.2 Key Concepts Principles of Gas Exchange Respiration is the sum of processes that move oxygen from air or water in the environment to all metabolically active tissues and move carbon dioxide from those tissues to the outside Oxygen levels are more stable in air than in water 39.3 Invertebrate Respiration Integumentary exchange • Some invertebrates that live in aquatic or damp environments have no respiratory organs; gases diffuse across the skin Gills • Filamentous respiratory organs that increase surface area for gas exchange in water Invertebrate Respiration Lungs • Saclike respiratory organs with branching tubes that deliver air to a respiratory surface Snails and slugs that spend some time on land have a lung instead of, or in addition to, gills Snails with Lungs Invertebrate Respiration Tracheal system • Insects and spiders with a hard integument have branching tracheal tubes that open to the surface through spiracles (no respiratory protein required) Book lungs • Some spiders also have thin sheets of respiratory tissue that exchange oxygen with a respiratory pigment (hemocyanin) in blood Insect Tracheal System trachea (tube inside body) spiracle (opening to body surface) Fig. 39-7, p. 685 A Spider’s Book Lung air-filled space blood-filled space book lung Fig. 39-8, p. 685 39.3 Key Concepts Gas Exchange in Invertebrates Gas exchange occurs across the body surface or gills of aquatic invertebrates In large invertebrates on land, it occurs across a moist, internal respiratory surface or at fluid-filled tips of branching tubes that extend from the surface to internal tissues 39.4 Vertebrate Respiration Fishes use gills to extract oxygen from water • Countercurrent flow aids exchange (blood flows through gills in opposite direction of water flow) Amphibians exchange gases across their skin, and at respiratory surfaces of paired lungs • Larvae have external gills Fish Gills gill cover Fig. 39-9a, p. 686 Fig. 39-9b, p. 686 mouth open gill cover closed Fig. 39-9b, p. 686 Fig. 39-9c, p. 686 mouth closed gill cover open Fig. 39-9c, p. 686 Countercurrent Flow gill filaments one gill arch water is sucked into mouth Water exits through gill slits A A bony fish with its gill cover removed. Water flows in through the mouth, flows over the gills, then exits through gill slits. Each gill has bony gill arches to which the gill filaments attach. Fig. 39-10a, p. 686 gill arch respiratory surface gill filament fold with a capillary bed inside water flow direction of blood flow oxygen-poor blood oxygenated blood from deep in body back toward body B Two gill arches with filaments C Countercurrent flow of water and blood Fig. 39-10 (b-c), p. 686 Animation: Bony fish respiration Frog Respiration A Lowering the floor of the mouth draws air inward through nostrils. B Closing nostrils and raising the floor of the mouth pushes air into lungs. C Rhythmically raising and lowering the floor of the mouth assists gas exchange. D Contracting chest muscles and raising the floor of the mouth forces air out of lungs, and the frog exhales. Fig. 39-11, p. 687 Animation: Frog respiration Vertebrate Respiration Reptiles, birds and mammals exchange gases through paired lungs, ventilated by chest muscles Birds have the most efficient vertebrate lungs • Air sacs allow oxygen-rich air to pass respiratory surfaces on both inhalation and exhalation Bird Respiratory System A Inhalation 1 Muscles expand chest cavity, drawing air in through nostrils. Some of the air flowing in through the trachea goes to lungs and some goes to posterior air sacs. B Exhalation 1 Anterior air sacs empty. Air from posterior air sacs moves into lungs. trachea anterior air sacs lung posterior air sacs C Inhalation 2 Air in lungs moves to anterior air sacs and is replaced by newly inhaled air. D Exhalation 2 Air in anterior air sacs moves out of the body and air from posterior sacs flows into the lungs. Fig. 39-12, p. 687 Fig. 39-12 (inset), p. 687 Animation: Bird respiration 39.5 Human Respiratory System The human respiratory system functions in gas exchange, sense of smell, voice production, body defenses, acid-base balance, and temperature regulation Airways Air enters through nose or mouth, flows through the pharynx (throat) and the larynx (voice box) • Vocal cords change the size of the glottis The epiglottis protects the trachea, which branches into two bronchi, one to each lung • Cilia and mucus-secreting cells clean airways Larynx: Vocal Cords and Glottis glottis closed vocal cords glottis open glottis (closed) epiglottis tongue’s base Fig. 39-14, p. 689 From Airways to Alveoli Inside each lung, bronchi branch into bronchioles that deliver air to alveoli Alveoli are small sacs, one cell thick, where gases are exchanged with pulmonary capillaries Muscles and Respiration Muscle movements change the volume of the thoracic cavity during breathing Diaphragm • A broad sheet of smooth muscle below the lungs • Separates the thoracic and abdominal cavities Intercostal muscles • Skeletal muscles between the ribs Functions of the Respiratory System Fig. 39-13a, p. 688 Oral Cavity (Mouth) Supplemental airway when breathing is labored Pleural Membrane Double-layer membrane with a fluid-filled space between layers; keeps lungs airtight and helps them stick to chest wall during breathing Intercostal Muscles At rib cage, skeletal muscles with roles in breathing. There are two sets of intercostal muscles (external and internal) Diaphragm Muscle sheet between the chest cavity and abdominal cavity with roles in breathing Nasal Cavity Chamber in which air is moistened, warmed, and filtered, and in which sounds resonate Pharynx (Throat) Airway connecting nasal cavity and mouth with larynx; enhances sounds; also connects with esophagus Epiglottis Closes off larynx during swallowing Larynx (Voice Box) Airway where sound is produced; closed off during swallowing Trachea (Windpipe) Airway connecting larynx with two bronchi that lead into the lungs Lung (One of a Pair) Lobed, elastic organ of breathing; enhances gas exchange between internal environment and outside air Bronchial Tree Increasingly branched airways starting with two bronchi and ending at air sacs (alveoli) of lung tissue Fig. 39-13a, p. 688 Fig. 39-13b, p. 688 bronchiole alveolar sac (sectioned) alveolar duct alveoli Fig. 39-13b, p. 688 Fig. 39-13c, p. 688 alveolar sac pulmonary capillary Fig. 39-13c, p. 688 Animation: Human respiratory system Animation: Examples of respiratory surfaces Animation: Vertebrate lungs