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,ffi, e,--L bI )uccesslon a - ataaaaalaaaaaf taraaaaaatattaaaaaaaaaatlaat i What llappened llere? i i ! ! : ! ! ! !a a a a a a o f. fh" two photographs at the right show the same area in Yellowstone National Park in Wyoming. Photograph A was taken soon after a major fire. Photograph B was taken a few years later. Observe the photographs carefully. 2. Make a list of all the differences you notice between the two scenes. a I i rtinh Posing a I a t a I a a over Questions How would you describe what happened during the time between the two photographs? What questions do you have about this process? a n I How are primary and secondary succession different? Reoding lip Before you read, write a definition of what you think the term succession might mean. As you read, revise your definition. 75'^ E 1988, a huge fire raged through Yellowstone National Park. The fire was so hot that it jumped from tree to tree without burning along the ground between them. In an instant, huge trees burst into flame from the intense heat. It took weeks for the fires to burn themselves out. All that remained of that part of the forest were thousands of blackened tree trunks sticking out of the ground like charred toothpicks. You might think it unlikely that Yellowstone could recover from such a disastrous fire. But within just a few months, signs of life had returned. First tiny green shoots of new grass appeared in the black ground. Then small tree seedlings began to grow again. The forest was coming back! Fires, floods, volcanoes, hurricanes, and other natural disasters can change communities in a very short period of time. But even without a disaster, communities change. The series of predictable changes that occur in a community over time is called succession. This section describes two types of succession: primary succession and secondary succession. Primary Succession Primary succession in an area where no ecosystem previously existed. Such an area might be a new island formed by the eruption of an undersea volcano, or an area of rock uncovered by a melting sheet of ice. You can follow the series of changes an area might undergo in Figure 26 below. These scenes show an area after aviolent volcanic eruption. At first there is no soil, just ash and rock. The first species to populate the area are called pioneer species. Pioneer species are often lichens and mosses carried to the area bywind or water. These species can grow on bare rocks with little or no soil. As these organisms grow they help break up the rocks. \Mhen they die, they provide nutrients that enrich the thin layer of soil that is forming on the rocks. Over time, plant seeds land in the new soil and begin to grow. The specific plants that grow depend on the biome of the area. For example, in a cool, northern area, eatly seedlings might include alder and cottonwood trees. As the soil grows older and richer, these trees might be replaced by spruce and hemlock. Eventually, succession may lead to a community of organisms that does not change unless the ecosystem is disturbed. Re4ching this stable community can take centuries. d eAaZAn*4t is the series of changes that occur What are some pioneer species? Figure 26 Primary succession occurs in an area where no ecosystem previously existed. A. After a volcanic eruption, the ground surface consists of ash and rock. B. The first organisms to appear are lichens and moss. C. Weeds and grasses take root in the thin layer of soil. D. Eventually, tree seedlings and shrubs sprout. Applying Concepts What determines the particular species thot oppear during succession? m,O -'" ;.j;i ,. :a**,r'l ,. J '** ** u,+,*ryo*.'.: Chopter2 Ea77 }i'rj is',$ .,ir *, .t .# i{r ,. {+r "its "s; nY,tsj;. fu'u *, lw *\;{ ,' Figure 27 Secondary succession occurs following a disturbance to an ecosystem, such as clearing a forest for farmland. When the farm is abandoned, the forest gradually returns. A. After two years, weeds and wildflowers fill the field. B. After five years, pine seedlings and other plants populate the field. C. After 30 years, a pine forest has grown up. D. After I00 years, a mixed forest of pine, oak, and hickory is developing in the field. Secondary Succession The changes following the Yellowstone fire were an example of secondary succession. Secondary succession is the series of changes that occur after a disturbance in an existing ecosystem. Natural disturbances that have this effect include fires, hurricanes, and tornadoes. Human activities, such as farming, logging, or mining, may also distuib an ecosystem. Unlike primary succession, secondary suc'cession occurs in a place where an ecosystem has previously existed. Secondary succession occurs somewhat more rapidly than primary succession. Consider, for example, an abandoned field in the southeastern United States. Follow the process of succession in such a field in Figure 27. Aft"er a century, a hardwood forest is developing. This forest is very stable and will remain for a long time. Of course, the particular species that come and go in the process of succession depend on the biome. 1. How are primary and secondary succession different? 2. What is a pioneer species? 3. Give two examples of natural disturbances and two examples of human disturbances that can result in secondary succession. 4. Thinking Critically Classifying Crass poking through the cracks in a sidewalk is an example of succession. ls this primary or secondary succession? Explain. 78.^ E 1E$gssffiapn;" I nerghborhood has itraiatuaffiffi dffi,.iffixT,g.H,?jji.