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Pigeonpea Botany and Production Practices Compiled by Faujdar Singh and D.L. Oswalt Skill Development Series no. 9 ICRISAT H u m a n Resource Development Program International Crops Research Institute for the Semi -Arid Tropics Patancheru, A n d h r a Pradesh 502 324, I n d i a 1992 Pigeonpea Botany and Production Practices Compiled by Faujdar Singh and D.L. Oswalt Skill Development Series no. 9 ICRISAT Human Resource Development Program International Crops Research Institute for the Semi-Arid Tropics Patancheru, Andhra Pradesh 502 324, India 1992 Human Resource Development Program P u b l i c a t i o n s i n t h i s S k i l l D e v e l o p m e n t S e r i e s (SDS) a r e i s s u e d s e m i f o r m a l l y for l i m i t e d d i s t r i b u t i o n t o p r o g r a m p a r t i c i p a n t s , colleagues, and collaborators. Copies may be requested by the SDS number. Constructive criticism from readers is w e l c o m e d b y : P r o g r a m Leader, Human Resource Development Program, ICRISAT. Acknowledgments Information has been taken from published and unpublished reports. This publication should not be cited as a reference. C o m m e n t s , suggestions, and encouragement from Dr Y . S . Chauhan, Dr D . M c D o n a l d , a n d D r Y . L . N e n e for c o m p i l i n g t h i s d o c u m e n t a r e gratefully acknowledged. T h a n k s t o M r S.V. P r a s a d R a o a n d M r P . Chenchaiah for computerizing this manuscript and to M r . M . M . B a b u for d r a w i n g f i g u r e s for c o m p u t e r s c a n n i n g . Contents Introduction 5 5 5 9 12 12 12 Origin and Distribution Pigeonpea Plant Inflorescence Flowering and Pollination Pod Development Seed and its Germination Production Practices 13 13 13 13 14 16 20 21 21 22 22 23 23 Climate Soils Land Preparation Responses to Nutrients Cropping Systems Sowing Time and Depth Methods of Sowing and Spacing Seed Rate and Seed Treatment Weed Management Irrigation Harvesting and Threshing Recording Observations MP 1. MP 2. D e t e r m i n a t i o n of S e e d R a t e Density for a Given Plant 24 Recording Morphological Characters 26 References 28 Evaluation 33 Introduction P i g e o n p e a [Cajanus cajan (L.) M i l l s p . ] b e l o n g s to t h e g e n u s Cajanus, s u b t r i b e Cajaninae, t r i b e P h a s e o l e a e , a n d f a m i l y a b a c e a e . It is also k n o w n a s r e d g r a m , t u r , a r h a r , g a n d u l (Spanish) a n d p o i s d ' A n g o l e (Sharma a n d G r e e n 1 9 8 0 ) , c o n g o b e a n ( E n g l i s h ) , p o i s d e C o n g o ( F r e n c h ) , a n d e r v i l b a d e C o n g o i n A n g o l a (Baldev 1 9 8 8 ) . Cajanus i s d e r i v e d f r o m a M a l a y w o r d ' k a t s c h a n g ' o r 'katjang' m e a n i n g p o d o r b e a n . Many s p e c i e s o f t h e c l o s e l y r e l a t e d g e n u s Atylosia s u c c e s s f u l l y c r o s s w i t h p i g e o n p e a (van d e r M a e s e n 1 9 8 0 ) . Origin and Distribution The pigeonpea name was first reported from plants used in Barbados. Once seed of this crop were considered very important there as pigeon f e e d (Plukenet 1 6 9 2 ) . Based on the range of genetic diversity of the c r o p i n I n d i a , V a v i l o v (1951) c o n c l u d e d t h a t p i g e o n p e a o r i g i n a t e d i n India. Several authors considered eastern Africa to be the center of o r i g i n o f p i g e o n p e a , a s i t o c c u r s t h e r e i n t h e w i l d f o r m (Zeven a n d Zhukovsky 1 9 7 5 ) . H o w e v e r , D e (1974) a n d V e r n o n R o y e s (1976) r e v i e w e d the pigeonpea origin and agreed in favor of India. Van der Maesen (1980) c o n c l u d e d t h a t I n d i a w a s t h e p r i m a r y c e n t e r o f o r i g i n a n d Africa was the secondary center of origin of pigeonpea. It is cultivated in many countries in the tropics and subtropics. Its drought tolerance and the ability to use residual moisture during the dry season m a k e it an important crop (Sheldrake 1 9 8 4 ) . P i g e o n p e a i s c u l t i v a t e d o n a b o u t 4.23 m i l l i o n h e c t a r e s i n t h e w o r l d , w i t h a n a v e r a g e a n n u a l p r o d u c t i o n o f 2.96 m i l l i o n t . Its average p r o d u c t i v i t y is 700 kg ha-1. The m a j o r pigeonpea growing area (3.58 m h a ) w a s in I n d i a w i t h a p r o d u c t i o n of 2.72 m t or a p r o d u c t i v i t y o f 7 6 0 k g h a - 1 (FAO 1 9 9 0 ; p e r s o n a l c o m m u n i c a t i o n b y ICRISAT Economics G r o u p ) . Africa, mainly Kenya and Uganda, accounts for 4% of the w o r l d ' s p r o d u c t i o n . The Caribbean and Central and South A m e r i c a p r o d u c e 2 % o f t h e t o t a l p i g e o n p e a o f t h e w o r l d (Sharma e t a l . 1981) . Pigeonpea Plant T h e p i g e o n p e a p l a n t i s e r e c t a n d b r a n c h i n g (Fig. 1 ) . The woody, leaves are trifoliate, and compound. It possesses taproot system. T h e p l a n t s g r o w i n t o w o o d y s h r u b s , 1-2 m annually harvested. It m a y a t t a i n a h e i g h t of 3-4 m w h e n perennial plant in fence rows or agroforestry p l o t s . stem is a strong tall when g r o w n as a Seedling W h e n s o w n u n d e r o p t i m a l m o i s t u r e a n d t e m p e r a t u r e (29°C-36°C), t h e s e e d testa splits open near t h e micropyle on the 2nd day. The tip of the r a d i c a l e l o n g a t e s a n d e m e r g e s f r o m t h e s e e d c o a t . O n t h e 3rd d a y t h e hypocotyl appears as an arch and continues to grow upward. The hypocotyl turns light purple. T h e s e e d l i n g e p i c o t y l e l o n g a t e s 3-7 c m b e f o r e t h e f i r s t t r i f o l i a t e l e a f e m e r g e s (Reddy 1 9 9 0 ) . HRDP SDS n o . 9 5 Figure 1. A stylized pigeonpea plant. Root Pigeonpea has a taproot system. The root growth, under ideal conditions, begins on the 2nd day after sowing of t h e seed. The testa splits open near the micropyle. The radical elongates and emerges from the seed coat (Chauhan, In p r e s s ) . The primary structure of the root i s t e t r a c h . Secondary thickening takes place as a result of c a m b i a l a c t i v i t y (Bisen a n d S h e l d r a k e 1 9 8 1 ) . The m a i n root of older plants becomes thick and woody. Pigeonpea roots are deep and wide spreading in the soil, with well-developed lateral roots. They may extend down more than 2 m, but extensive development takes place in the upper 60 cm of t h e soil p r o f i l e (Sheldrake and N a r a y a n a n 1 9 7 9 ; Natarajan and Willey 1 9 8 0 ) . Short-duration genotypes develop a smaller root system t h a n long-duration genotypes (Sheldrake and Narayanan 1 9 7 9 ) . Lateral roots were found much longer in spreading t y p e s t h a n i n t h e e r e c t t y p e s (Mahta a n d D a v e 1 9 3 1 ) . 6 HRDP SDS no. 9 Modules Pigeonpea is nodulated by the cowpea group of rhizobia, mainly on the u p p e r 30 cm of the root system. Nodulation starts approximately 15 d a y s a f t e r s o w i n g (DAS) a n d c o n t i n u e s u p t o 120 d a y s . It declines t o w a r d s p o d f i l l i n g (Kumar R a o 1 9 9 0 ) . The nodule development is through the meristematic zone, arching around the apical end and the medulla contains many bacteroid-filled cells. S o m e t i m e s t h e l a t t e r a r e h i g h l y v a c u o l a t e d (Bisen a n d Sheldrake 1 9 8 1 ) . T h e n o d u l e s d i f f e r i n s i z e f r o m 2-4 m m . T h e y m a y b e spherical, oval, elongate, or branched. Stem The pigeonpea has a strong woody stem. The primary vascular tissue of the stem is organized into strands connecting the nodes; each strand is associated with a ridge on the stem. Collenchymatous bundle caps u n d e r l i e t h e e p i d e r m i s o f t h e r i d g e s (Bisen a n d S h e l d r a k e 1 9 8 1 ) . During the vegetative phase, the xylem parenchyma, and the medullary rays contain starch. This disappears in the reproductive p h a s e , s h o w i n g t h a t t h e r e s e r v e s a r e m o b i l i z e d for p o d d e v e l o p m e n t (Sheldrake 1 9 8 4 ) . There are cells within the phloem region and the outer part of the pith containing densely staining material. These cells are joined end to end, forming d u c t s . Such secondary ducts differentiate at an early stage in the primary tissue and are also formed within the secondary phloem. They occur in the stem, p o d s , and roots. The plant's damaged tissues secrete droplets of a clear fluid exudate from the secondary d u c t s o n t o t h e cut s u r f a c e . T h i s t u r n s red o n e x p o s u r e t o a i r a n d covers the wound with a reddish varnish-like material. This exudate has a stringent taste and may have a role in protection against pests (Sheldrake 1 9 8 4 ) . Branches The branching pattern in pigeonpea depends on genotype and spacing between rows and plants. At a w i d e s p a c i n g , it m a y f o r m a b u s h a n d at narrow spacing it may remain compact and upright. For agronomic purposes pigeonpea plants can be grouped as compact (erect), semispreading (semierect), and spreading types. Based on the f l o w e r i n g p a t t e r n i t m a y b e d e t e r m i n a t e o r n o n d e t e r m i n a t e (Fig. 2 ) . The determinate type completes the vegetative phase and then enters into the reproductive p h a s e . In this type, the apical bud of the main shoot develops into an inflorescence, and the sequence of inflorescence p r o d u c t i o n is b a s i p e t a l (developing in the direction of base). The nondeterminate type shows continuous vegetative and reproductive phases. In this type, the flowering starts at nodes behind the apex and proceeds both acropetally and basipetally. Another group is semideterminate between the determinate and nondeterminate types. It includes late-maturing genotypes where branching starts from different angles, but most of the pods are at the upper region of the plant. HRDP SDS no. 9 7 Figure 2. Branching pattern in pigeonpea. Leaves The first two leaves in the seedling called primary leaves, are s i m p l e , o p p o s i t e , a n d c a d u c e u s (Fig. 3 ) . The later leaves are pinnately trifoliate with lanceolate to elliptical leaflets that are acute at both ends and are spirally arranged. The leaflets are borne on a rachis, which is swollen at the base (pulvinus). The leaf size v a r i e s f r o m 6-17 c m i n l e n g t h a n d a r e a b o u t t h e s a m e w i d t h . The rachis v a r i e s f r o m 2-4 c m a n d t h e t e r m i n a l l e a f l e t s a r e 4-8 c m b y 2-3.5 cm. The lateral leaflets are slightly smaller. There is genetic variability in the size, shape, and color of leaves. The leaves are pubescent with more on the lower than the upper surface. The hair types are simple or g l a n d u l a r . The latter are s p h e r i c a l a n d c o n t a i n a y e l l o w o i l y m a t e r i a l , p r o b a b l y r e s p o n s i b l e for t h e f r a g r a n c e o f p i g e o n p e a p l a n t s (Bisen a n d S h e l d r a k e 1 9 8 1 ) . 8 HRDP SDS no. 9 Figure 3. Pigeonpea seedling showing primary and trifoliate leaves. (Source: R e d d y 1990) Inflorescence In most cultivars, flowers are borne on terminal or auxiliary racemes (4-12 cm) a n d a r e c a r r i e d o n a l o n g p e d u n c l e (Fig. 4 a ) . The raceme i n f l o r e s c e n c e f o r m a t e r m i n a l p a n i c l e in n o n d e t e r m i n a t e t y p e s a n d a somewhat corymb-shape bunch in the determinate types. These are grouped together at the end of branches in late types and distributed a l o n g t h e b r a n c h e s i n e a r l y , m e d i u m , a n d i n d e t e r m i n a t e t y p e s (Sharma and Green 1 9 8 0 ) . T h e n u m b e r o f r a c e m e s plant" 1 i n t h e p i g e o n p e a w o r l d c o l l e c t i o n s r a n g e d f r o m 6 t o 915 ( R e m a n a n d a n e t a l . 1 9 8 8 ) . F l o w e r i n g p r o c e e d s a c r o p e t a l l y (in t h e d i r e c t i o n o f a p e x ) b o t h w i t h i n t h e r a c e m e and on the branch. Flower T h e f l o w e r s (Fig. 4a) a r e c l u s t e r e d a t t h e t o p o f t h e p e d u n c l e . The p e d u n c l e s a r e 1-8 c m l o n g . Flowers are frequently yellow. The bracts (Fig. 4d) a r e s m a l l w i t h a t h i c k m i d d l e n e r v e . They are ovate-lanceolate with hairy margins and curved inwards to form a boat l i k e s t r u c t u r e t o e n c l o s e o n e t o t h r e e y o u n g l a t e r a l b u d s (Fig. 4 b ) . T h e p e d i c e l i s t h i n , 5-15 m m l o n g , a n d c o v e r e d w i t h h a i r . T h e f l o w e r s a r e m o s t l y y e l l o w (Fig. 4c) a n d p a p i l e o n a c e o u s o r completely b i s e x u a l and zygomorphic (Sundaraj and Thulasidas 1 9 8 0 ) . The color of the flowers vary from yellow to yellow with purple veins HRDP SDS no. 9 9 and from yellow to diffused red. length. The flowers are usually 2 cm in Calyx The calyx is gamosepalous with five lobes. The calyx tube is campanulate (bell-shaped) with nerved teeth. The upper two teeth are subconnate. T h e l o w e r t h r e e a r e f r e e a n d s p r e a d i n g (Fig. 4 e ) . The upper lobes are paired, free or partly free, and the lower one is the longest. d b Figure 4. e C a Pigeonpea inflorescence (a), bract (d), and calyx ( e ) . ( S o u r c e : R e d d y 1990) bud (b), flower (c), Corolla The corolla is zygomorphic (yoke-shaped flowers symmetrical about one plane) and bright yellow. The petals are imbricate and of three prominent types; the standard, wings, and a keel. The standard is b r o a d , l a r g e , a u r i c l e d , a n d e r e c t (Fig. 5 a ) . The wings are obliquely o b o v a t e w i t h a n i n c u r v e d c l a w (Fig. 5 b ) . The keel petals are obtuse ( r o u n d ) , i n c u r v e d (Fig. 5 f ) , a n d b o a t s h a p e d (Fig. 5 c ) . The keel c o v e r s t h e a n d r o e c i u m (stamens) a n d g y n a e c e u m (female o r g a n s ) o f t h e flower. Normally the standard and wings are bright yellow. The keel is greenish yellow. Aestivation is a descending imbricate or one whorl outside is free and the one inside has both margins overlapped. The other whorls overlap by only one margin. 10 HRDF SDS no. 9 Androecium S t a m e n s a r e 1 0 , d i a d e l p h o u s (in t w o b u n d l e s 9+1) w i t h 9 f u s e d in a c o l u m n a n d 1 f r e e (Fig. 5 d ) . T h e f r e e s t a m e n f i l a m e n t (4-7 m m ) i s attached at the base. The other filaments are fused t o g e t h e r for t h e greater part and enclose the gynaeceum. The upper free p o r t i o n of the filaments terminate in anthers. T h e a n t h e r s a r e u n i f o r m , a b o u t 1-mm long. The two halves of the anthers are joined by a relatively large, s t e r i l e c o n n e c t i v e t u b e t h a t i s b a s i f i x e d (Fig. 5 e ) . The anthers are light or dark yellow, dorsifixed. Of the 10 stamens, four have short filaments and six, including a posterior one, have long filaments. The short anthers have blunt lobes and the long ones pointed lobes. The p o l l e n p r o d u c e d by short stamens is generally used for self f e r t i l i z a t i o n (Bahadur e t a l . 1 9 8 1 ) . Gynaeceum The ovary is superior, subsessile, flattened dorsoventrally w i t h a l o n g s t y l e (Fig. 5 f ) . It has a very short stalk, densely pubescent a n d g l a n d u l a r p u n c t a t e (dotted o r p i t t e d ) w i t h t w o t o n i n e o v u l e s , marginal placentation, monocarpellary, and unilocular. The style is long, filiform, upturned beyond the middle region, and glabrous. It i s a t t a c h e d t o a t h i c k e n e d , i n c u r v e d , a n d c a p i t a t e (swollen) s t i g m a (Fig. 5 g ) . e d a b C Figure 5. g f Pigeonpea corolla: standard (a), wings keel (c), androecium (d), anther (e), gynaecium (f), and stigma ( g ) . ( S o u r c e s : R e d d y 1 9 9 0 ; a n d B a l d e v 1988) HRDP SDS no. 9 (b), 11 Flowering and Pollination P i g e o n p e a is a s h o r t - d a y p l a n t , t h o u g h a c o m b i n a t i o n of p h o t o p e r i o d and temperature can affect the time of flowering and maturity of p i g e o n p e a g e n o t y p e s a c c o r d i n g t o d a t e o f s o w i n g (Green e t a l . 1 9 7 9 ) . In a fully d e v e l o p e d b u d , anthers surround the stigma and dehisce a day before the flower o p e n s . Anthesis in pigeonpea starts from 0600 and continues till 1600. The peak anthesis period recorded at ICRISAT i s b e t w e e n 0 9 0 0 a n d 1 0 0 0 (Sharma a n d G r e e n 1 9 8 0 ) . The duration of flower opening depends on the climate and environment. This varies from 6 to 36 h. F e r t i l i z a t i o n occurs on the day of p o l l i n a t i o n . Pigeonpea is an often cross-pollinated crop. The extent of c r o s s p o l l i n a t i o n r a n g e s f r o m 3 t o 4 0 % (Khan 1 9 7 3 ) , w i t h a n a v e r a g e o f 2 0 % . A p l a n t p r o d u c e s m a n y f l o w e r s o f w h i c h o n l y 1 0 % set p o d s (Pathak 1970). Pod Development The pedicel of the flower contains small vascular bundles surrounded by a ring of fibers. D u r i n g t h e 1st w e e k o f a n t h e s i s , t h e e n d o s p e r m undergoes rapid development. The nuclei take up a parietal position, forming a large vacuole in the center of the embryo sac. The embryo sac elongates at the chalazal region and forms a haustorium. The haustorium penetrates into the nucellar tissue. This is instrumental in absorbing food material that is used by the developing embryo. By the end of the 2nd week there are still large amounts of endospermous tissues and within the embryo, distinct cotyledons are seen. Further development of the seed involves rapid growth of the cotyledons, and almost complete degeneration of the endosperm. In the cotyledons, synthesis of starch and protein starts about 17 days after pollination a n d c o n t i n u e s f o r 14 d a y s (Sehgal et a l . 1 9 8 7 ) . In e a c h r a c e m e 1 to 5 pods may mature, and rarely up to 10. Pods are of various color; green, purple, dark purple, or mixed green and purple. Pods with deep constrictions in shape are beaded, while others are somewhat flat. The seeds per p o d range from two to seven, and sometimes up to nine. The seeds are in separate locules a n d t h e c r o s s w a l l s d e v e l o p d u r i n g t h e 1st w e e k a f t e r f e r t i l i z a t i o n . The pod wall develops more rapidly than the young seeds. Seed development is v i s i b l e 7 days after p o l l i n a t i o n . A p o d is formed 15-20 days after fertilization. Seeds reach physiological maturity in 30 days and are ready for harvest at a lower m o i s t u r e content in 40 d a y s (Rao a n d R a o 1 9 7 4 ) . There is little or no shattering of mature pods in the field. Seed and its Germination The color of the seed ranges from silver, white, cream, fawn, black, pink, or red to purple. They are blotched or speckled. Pigeonpea 1 0 0 - s e e d m a s s r a n g e s f r o m 2.8 t o 2 2 . 4 g w i t h t h e c u l t i v a t e d v a r i e t i e s r a n g i n g f r o m 7.0 t o 9.5 g . Seed shapes are oval, pea-shaped, square, or elongate. The most common is a pea-shaped seed found in l a r g e seeded late varieties. The number of seeds p o d - 1 ranges from 2 to 8. T h e c u l t i v a t e d g e n o t y p e s p o s s e s s 3-4 s e e d s p o d - 1 (Reddy 1 9 9 0 ) . The cotyledons are yellow. Germination is hypogeal and there is no known dormancy. The seedlings emerge from depths of up to 5 cm. 12 HRDP SDS no. 9 Production Practices Climate Pigeonpea is predominantly a crop of tropical areas mainly cultivated in semi-arid regions of India and Kenya. It is also cultivated in subhumid regions of Uganda, the West Indies, Myanmar, and the Caribbean regions. R e d d y a n d V i r m a n i (1981) c o n c l u d e d t h a t p i g e o n p e a c a n b e g r o w n b e t w e e n 14°N a n d 28°N l a t i t u d e , w i t h a t e m p e r a t u r e r a n g i n g f r o m 26° to 30°C in t h e r a i n y s e a s o n (June to O c t o b e r ) a n d 17° to 22°C i n t h e p o s t r a i n y (November t o M a r c h ) s e a s o n . The amount of daily g l o b a l s o l a r r a d i a t i o n v a r i e s f r o m 4 0 0 t o 4 3 0 cal cm - 2 d a y - 1 i n t h e rainy season and 380-430 cal cm-2 day-1 in the postrainy season. Mean annual rainfall ranges from 600 to 1400 mm, of which 8 0 % - 9 0 % is . received in the rainy season. The length of growing season extends from 120 to 180 d a y s . Isoclimes similar to India exist in western A f r i c a a n d s o u t h e r n S u d a n , a s u i t a b l e e n v i r o n m e n t for g r o w i n g pigeonpea. Pigeonpea is very sensitive to low radiation at pod development, therefore flowering during the monsoon and cloudy weather, leads to poor pod formation. Soils In India, pigeonpea is cultivated on Entisols, Alfisols, Enceptisols, and Vertisols. The E n t i s o l s found in the a l l u v i a l - soil belt of the I n d o - G a n g e t i c r e g i o n a r e d e e p l o a m s , s l i g h t l y a l k a l i n e (pH 7 . 5 - 8 . 5 ) , with about 150-200 mm available water storage capacity in 2 m of soil. The V e r t i s o l s are characterized by 4 0 - 6 0 % clay in the surface soil h o r i z o n s , p H a r o u n d 8.0 w i t h a water holding capacity between 150-300 m m , a n d t h e a v a i l a b l e w a t e r i n t h e t o p 1.5-2.0 m o f s o i l . Alfisols a r e n e u t r a l in r e a c t i o n (pH 6.5-7.0) a n d r e l a t i v e l y s h a l l o w w i t h a low-clay content. T h e y a r e o f t e n s a n d y l o a m a n d c a n r e t a i n a b o u t 100 m m a v a i l a b l e w a t e r i n t h e r o o t p r o f i l e (Reddy a n d V i r m a n i 1 9 8 1 ) . Pigeonpea, being sensitive to water logging, requires a well-drained soil. It does not grow well in saline soil, but can withstand drought reasonably well. Responses to lime indicated by increase in shoot g r o w t h a n d n o d u l a t i o n , h a v e b e e n r e p o r t e d i n s o i l s w i t h p H b e l o w 5.0 (Edwards 1 9 8 1 ) . Land Preparation L a n d p r e p a r a t i o n for p i g e o n p e a r e q u i r e s a t l e a s t o n e p l o w i n g d u r i n g the dry season followed by 2 or 3 h a r r o w i n g s . The "summer" plowing h e l p s i n m i n i m i z i n g t h e w e e d f l o r a a n d t o c o n s e r v e m o i s t u r e (Chandra et al. 1 9 8 3 ) . W e l l - d r a i n e d s o i l s a r e n e c e s s a r y for g o o d root a n d nodule development. Contour beds or a ridge-and-furrow system are useful in preventing water logging by draining excess surface water, and in preventing soil erosion. O r g a n i c m a n u r e m a y b e a p p l i e d 2-4 w e e k s b e f o r e s o w i n g . I n a c i d i c s o i l s 2-4 t ha - 1 of l i m e is i n c o r p o r a t e d 3-4 w e e k s b e f o r e s o w i n g to neutralize the acidity. In light soils, a basal application of aldrin 5% d u s t @ 30 kg ha - 1 p r e v e n t s t e r m i t e i n f e s t a t i o n (Chandra et a l . 1983). HRDP SDS no. 9 13 Responses to Nutrients Responses to general meager. Nitrogen fertilizer applications in pigeonpea are quite (N) K u l k a r n i a n d P a n w a r (1981) r e v i e w e d t h e s t u d i e s i n I n d i a o n t h e pigeonpea response to N. They concluded the effect was almost negligible. However, in some situations a starter application of 2 0 - 2 5 kg N ha - 1 w a s b e n e f i c i a l . Rhizobium inoculation R e s p o n s e s t o Rhizobium i n o c u l a t i o n h a v e b e e n i n c o n s i s t e n t (Panwar a n d Misra 1973; Panwar 1 9 7 5 ) . Increases in grain yield of pigeonpea i n o c u l a t e d w i t h e f f e c t i v e Rhizobium r a n g e d f r o m 1 9 t o 6 8 % (Kumar R a o 1990). Phosphorus (P) Responses to phosphorus applications have been positive and in some c a s e s h i g h l y s i g n i f i c a n t i n p i g e o n p e a (Pathak 1 9 7 0 ) . K u l k a r n i a n d P a n w a r (1981) s u m m a r i z e d t h e p h o s p h o r u s r e s p o n s e s t u d i e s i n I n d i a . They concluded that applications of 17-26 kg P ha-1 increased seed yield by 300-600 kg ha-1. Responses of pigeonpea to P fertilization would not be expected in soils w i t h m o r e than 5 mg kg-1 extractable P by Olsens bicarbonate extraction; although lower values would not necessarily p r e d i c t a response (Johansen 1 9 9 0 ) . Potassium (K) Pigeonpea does not respond to potassium chloride applications unless i t i s g r o w n o n l o w a v a i l a b l e p o t a s s i u m s o i l s (Kulkarni a n d P a n w a r 1981). Zinc (Zn) Most of t h e p i g e o n p e a cultivars are susceptible to zinc deficiency. T h e r e f o r e , f o l i a r a p p l i c a t i o n s o f 2-4 p p m z i n c a s 0 . 5 % z i n c s u l f a t e with 0.25% lime have b e e n effective to over-come zinc deficiencies (Saxena a n d S i n g h 1 9 7 0 ) . Nutrient deficiency symptoms The deficiency symptoms of m a c r o 2) are listed. 14 HRDP (Table 1 ) and micro-nutrients SDS no. 9 (Table Table 1. Deficiency symptoms of major nutrients in pigeonpea. Nutrient Deficiency symptoms on plant Nitrogen (N) Phosphorus Potassium Calcium Sulfur Young leaves uniformly become pale-green, greenish-yellow or paleyellow. Old leaves show an interveinal chlorosis and only veins remain green, leaves soon abscise (shed). (P) (K) Leaf tips turn yellow or brown. Yellowing spreads from the tip outward along the margin and may coalesce with similar yellow areas at extremities of the lateral veins. The leaf tip becomes scorched as symptoms become severe. The scorching may spread around the leaf margin. There is a yellow band between the scorched area and healthy-green tissues in the early stages. The affected leaves not showing these symptoms are generally dark-green and plants are stunted. (Ca) Light-green patches, irregular in outline, appear randomly in interveinal areas or around the leaf margin. Patches may turn brown, particularly on the young and severely affected leaves; such leaves abscise. Stems are weak and prostrate. (S) Magnesium Even yellowing across lamina of mainly young leaves. leaves become yellow. (Mg) (Sources: Table 2. Difficult to diagnose, it delays flowering. In general plants are stunted due to failure of internode elongation. The foliage remains dark-green and old leaves shed. Eventually old Interveinal Beginning in older leaves, in-rolling of leaf margins. regions light-green. Mesophyll of interveinal regions become lightgreen, leading to eventual necrosis. N i c h o l s 1964 and 1 9 6 5 ; J o h a n s e n 1990) Deficiency symptoms of micro nutrients in pigeonpea. Nutrient Deficiency symptoms on plant Zinc (Zn) Stunted growth, narrowing of leaves with pale-green or yellow appearance. Interveinal chlorosis starting from tip of leaflets and spreading to the remaining area leaving only the midrib green. Curling and shedding of leaflet when it becomes two-thirds chlorotic. Iron (Fe) Interveinal areas of young leaves become pale-green, veins and adjacent tissues remain dark-green. In severe cases, the entire area of the leaflet becomes chlorotic with small necrotic patches. Manganese Boron (Mn) (B) Molybdenum (Source: Fading of the green lamina of the middle leaves without affecting the veins. The small white and brown spots, first appearing in the chlorotic areas, coalesce and form brown necrotic lesions. It reduces size, number and area of leaflets, and delays flowering by arresting the growth of apical shoots. Boron helps in division and elongation of cells. Its deficiency results in dieback, rosetting, multiple branching, and death of seedlings. (Mo) Nitrogen fixation requires molybdenum for incorporation of the nitrogenase enzyme. Like nitrogen deficiency it causes general yellowing of the plant. K a t y a l 1 9 8 1 ; N i c h o l s 1964 a n d 1 9 6 5 ) HRDP SDS no. 9 15 The n u t r i e n t s required for selected levels are given in Table 3. of grain yield in pigeonpea Table 3. levels of ha-1) Estimated nutrients required grain yield of pigeonpea. (kg to produce selected Grain yield (kg ha -1 ) Nutrients N P K Ca Mg Mn Zn Cu (Source: 500 132 10 31 38 27 0.10 0.16 0.07 1000 1500 148 11 36 39 28 0.11 0.19 0.07 Calculations 164 13 41 40 29 0.11 0.22 0.07 based on 2000 181 15 47 41 29 0.12 0.25 0.08 Dalai 2500 3000 197 17 52 42 30 0.12 0.28 0.09 213 18 58 43 31 0.13 0.31 0.09 3500 230 20 63 44 32 0.13 0.35 0.10 4000 246 22 69 45 33 0.13 0.38 0.11 1980) Economic level of fertilization Pigeonpea cultivated in the arid and the semi-arid tropics is high-risk crop. The farmers are reluctant to use even major nutrients with proven responses to nutrient application (Chaudhary and Bhatia 1971; Dalai and Quilt 1 9 7 7 ) . a K u l k a r n i a n d P a n w a r (1981) c o n c l u d e d t h a t a n a p p l i c a t i o n o f 20 kg N, 17 kg P, a n d 17 kg K ha-1 c o u l d g i v e a s a t i s f a c t o r y yield on medium-black, red, and mixed black and red soils. This gave a cost-benefit ratio of R s . 1.28-4.07 Rupee-1 cost of fertilizer. However, in red and laterite soils 20-17-17 kg ha-1 gave a cost-benefit ratio of R s . 6.35-RS.8.80 Rupee-1 cost of f e r t i l i z e r (ICAR 1 9 7 9 ) . I n I n d i a , d i a m m o n i u m p h o s p h a t e (DAP) i s the most commonly used fertilizer for pigeonpea. This is used at a rate of 100 kg ha-1. It s u p p l i e s about 18-20-0 (Tandon 1 9 8 7 ) . At I C R I S A T , 10 kg ha-1 N ( a m m o n i u m s u l f a t e ) a n d 17 kg P ha-1 (single super p h o s p h a t e ) are applied in many e x p e r i m e n t s . The single super phosphate has the advantage of containing sulfur (S) . Cropping Systems A cropping system is defined as a combination of crops in space and time. The objective of any given system should be to provide the farmer a high and sustainable level of returns. In agronomic terms, the system makes efficient use of the basic resources necessary for plant growth (Willey et a l . 1 9 8 1 ) . Pigeonpea is grown both under sole and intercropping systems. 16 HRDP SDS n o . 9 Sole cropping Early-maturing (100-120 days) genotypes are grown as a sole crop. These genotypes have a higher harvest index with an average of 3 4 % compared to medium-maturing genotypes at 2 4 % (Sheldrake and Narayanan 1979). Therefore, from the cropping system point of view, early-maturing genotypes are better able to 'complement' by p r o v i d i n g an i n c r e a s e d opportunity for a second crop (Willey et al. 1981). Postrainy season pigeonpea are also cultivated as a sole crop. This avoids the wet conditions associated with the rainy season, gives less incidence of pests and diseases, and makes better use of residual soil moisture. Moreover, postrainy season sole crops were found more efficient than the rainy season sole crops (Willey et a l . 1 9 8 1 ) . Pigeonpea intercropping Pigeonpea is commonly intercropped with a wide range of crops. India, it was estimated that 80%-90% of the pigeonpea were i n t e r c r o p p e d (Aiyer 1 9 4 9 ) . W i l l e y e t a l . (1981) g r o u p e d pigeonpea intercrops into three broad categories: a) With cereals (sorghum, m a i z e , p e a r l m i l l e t , finger millet, and rainfed r i c e ) . setaria, b) With legumes soybean, and black (groundnut, cowpea, phaseolus b e a n ) . c) With long-season cassava). annuals mung bean, (caster, cotton, gram, sugarcane, and Chandra et al. (1983) l i s t e d h i g h l y p r o d u c t i v e i n t e r c r o p s p i g e o n p e a for d i f f e r e n t r e g i o n s i n India (Table 4 ) . Table Inter 4. Important intercrops Sowing crop Ratio of component pattern P a i r e d rows at 3 0 : 3 0 : 6 0 cm 2:1 Pearl millet P a i r e d r o w s at 3 0 : 3 0 : 6 0 cm 2:1 P a i r e d rows at 4 0 : 4 0 : 8 0 cm or u n i f o r m rows at 60 cm 2:1 Pigeonpea + upland rice U n i f o r m rows at 60-75 cm 2:2 Legumes Pigeonpea U n i f o r m rows at 75-90 cm 2:2 Maize + pigeonpea + groundnut Pigeonpea + soybean, mung bean or black gram U n i f o r m rows at 75 cm U n i f o r m rows at 50 cm HRDP of for pigeonpea. Cereals Sorghum + pigeonpea + pigeonoea In or SDS no. 9 crops 2:2 2:1 17 The advantages of intercrops are (Willey et al. 1981): a. Intercropping confers greater yield stability than sole cropping. For instance, if one crop fails or grows poorly, the other crop can produce to some extent. b. Under stress conditions, intercropping depression than sole cropping. c. Intercropping may reduce the incidence of w e e d s . The poor canopy cover and slow growth of pigeonpea in the early stages makes a sole crop susceptible to weed. Therefore, a fast-growing intercrop not -only gives a potential additional yield benefit, but also reduces the need for weeding (Shetty and Rao 1 9 7 9 ) . gives less yield Varieties Some of the important varieties of pigeonpea released in India for the different maturity groups (Tables 5a, 5b, 5c, and 5d). Table 5a. Extra-short-duration varieties o f pigeonpea released in India. Variety Pedigree Y e a r of release (State) Time to maturity (days) UPAS-120 Selection from P 4758 1976 (UP) 120-140 Nondeterminate, y i e l d 1.5-1.8 t h a - 1 . Pant - 1975 (UP) 120-130 Semierect, determinate, dwarf 3-4 s e e d s pod - 1 , 8 g (100 s e e d ) - 1 . Prabhat TI X T 190 1975 (UP) 110-120 D e t e r m i n a t e , d w a r f (100-120 c m ) , clustered fruiting, yield 1.2-1.5 t h a - 1 . ICPL 87 (Pragati) T21 x JA 277 1986 (Penin. India) 120-130 D e t e r m i n a t e , 10.5 g (100 brown seeded, potential y i e l d 2.5-3.0 t h a - 1 . ICPL 151 (Jagriti) ICP 6997 x Prabhat 1990 (NW India) 120-130 D e t e r m i n a t e , 10.5 g (100 s e e d ) - 1 , cream color, potential yield 2.5-3.0 t h a - 1 . C02 Selection from P 4728 1977 (Tamil Nadu) 120-130 B r o w n s e e d e d , 7 g (100 s e e d ) - 1 , a v e r a g e y i e l d 1.2-1.5 t ha -1 Mass selection from P 8 - 9 1981 130-140 (Punjab) AL 18 15 A3 HRDP Characteristics seed)-1, S m a l l s e e d e d 6-8 g (100 s e e d ) - 1 , p o t e n t i a l y i e l d 2.0 t ha - 1 , a v e r a g e y i e l d 1.4 t h a - 1 . SDS no. 9 Table 5b. Short-duration varieties o f pigeonpea ralaasad i n India. Variety Pedigree Pusa Brazil T190 Ageti 1-1 x Y e a r of release (State) Time to Maturity (days) Characteristics 1971 150-160 Dwarf compact plant, determinate, b o l d s e e d e d 9 g (100 s e e d ) - 1 , 4-5 s e e d s pod - 1 , p o t e n t i a l y i e l d 2.5 t h a - 1 . T21 TI x T190 1961 (UP) 150-170 S e m i s p r e a d i n g , loose p l a n t a n d indeterminate branching. Small seeded, 7 g (100 s e e d ) - 1 . HY2 PI 4628 1978 (AP) 140-150 S e m i e r e c t t a l l , p u r p l e stem, w h i t e seed, b o l d , 11-12 g (100 s e e d ) - 1 , p o t e n t i a l y i e l d 2.5-2.7 t h a - 1 , a v e r a g e 1.3-1.9 t h a - 1 . Pusa 84 Pusa Ageti T21 C01 Local (Tamil Table 5c. x Nadu) 140-150 1980 (N zone) 1970 (TN) 135-140 Medium tall, semispreading, pod - 1 , 7.5 g (100 s e e d ) - 1 , brown seeded. 3-4 Photoinsensitive, brown seeded, 7 g (100 s e e d ) - 1 , a v e r a g e y i e l d 1.5 t h a - 1 . Medium-duration varieties o f pigeonpea developed i n India. Variety Pedigree Year of release (State) Time to maturity (days) HY 1 PI 3704 1975 (AP) 160-170 S p r e a d i n g , p u r p l e stem, w h i t e seed, 9-10 g (100 s e e d ) - 1 , p o t e n t i a l yield 1.9-2 t ha - 1 , a v e r a g e 1.2-1.5 t h a - 1 . HY PI 2817-1A 1980 (AP) 160-170 E r e c t t a l l p l a n t (230-240 c m ) , g r e e n stem, w h i t e b o l d seeded, 18-20 g (100 s e e d ) - 1 , 4-5 seeds pod - 1 , p o t e n t i a l y i e l d 3.0-4.0 t h a - 1 , a v e r a g e yield 1.6-2.0 t h a - 1 . PI 3701 1980 (AP) 160-170 S e m i e r e c t , p u r p l e stem, m e d i u m h e i g h t (170 c m ) , b r o w n seeded, 12.9 g (100 s e e d ) - 1 , 4-5 seeds pod - 1 . B r a z i l 1X N P W R 15 1971 (Bihar) 180-200 M e d i u m t a l l (200-220 c m ) , s e m i s p r e a d i n g , p r o f u s e b r a n c h i n g , 4-5 seeds pod - 1 , b r o w n b o l d s e e d s , 10 g (100 s e e d ) - 1 ' wilt r e s i s t a n t , yield 3.0 t h a - 1 , a v e r a g e y i e l d 1.65 t ha - 1 . Local selection 1981 (MP) 165-195 M e d i u m tall (172-194 c m ) , 3-4 seeds pod - 1 , 10 g (100 s e e d ) - 1 , p o t e n t i a l y i e l d 3.0 t ha - 1 , a v e . 1.65 t ha - 1 . Local selection 1978 160-179 (Maharashtra) S e m i s p r e a d i n g , wilt resistant height (200-220 c m ) , 9-10 g (100 seed) - 1 Kanke 1976 (West Bengal) S e m i c o m p a c t , 11 g (100 s e e d ) - 1 . P o t e n t i a l y i e l d 2.5 t ha - 1 , a v e r a g e y i e l d 1.7 t ha - 1 . 3A HY 5 Mukta AS BDN S20 (R60) 71-37 1 1XB7 HRDP 180-190 Characteristiscs SDS no. 9 19 Table 5d. Late-duration varieties o f pigaonpaa ralaaaad i n India. Variety Pedigree C11 Local selection (AP) Bahar D1258 Local X Laxmi (Kanke-3) Gwalior 3 Time to Characteristic maturity (days) 200-220 Medium tall, profuse branching, s p r e a d i n g , b r o w n s e e d , 8.5 g (100 s e e d ) - 1 . 1973 (Bihar) 220-240 Compact, semierect plant, brown round s e l e c t i o n 4-5 s e e d s pod - 1 , y i e l d p o t e n t i a l 3 t h a - 1 , a v e r a g e y i e l d 2.25 t h a - 1 . BR-183X Local 1974 (West Bengal) 180-220 Perennial, semispreading, field t o l e r a n t t o w i l t , potential yield 2 t ha-1, a v e r a g e y i e l d 1.75 t h a - 1 . Local selection 1960/ 270> 1 9 8 0 (MP) (AP = A n d h r a P r a d e s h ; Pradesh) (Source Year of release (State) for Tables T a l l 2 5 0 - 3 0 0 cm, s p r e a d i n g , light b r o w n s e e d , 7-8 g (100 s e e d ) - 1 . MP = Madhya Pradesh; 5a, 5b, 5c, and TN = Tamil Nadu; 5d: Chandra et UP = Uttar al. 1983) Sowing Time and Depth In the rainfed and dry areas pigeonpea are sown with the onset of the monsoon. Earlier sowing gives higher yields in India. When sowing e x t r a - e a r l y and e a r l y - m a t u r i n g v a r i e t i e s in t h e 1st fortnight of June, the field is available for postrainy season crops by the end of November. Therefore, sowing should not be delayed beyond June. The sowing of medium and late-maturing varieties, under rainfed conditions, should be done during June or July at the onset of the monsoon. This should be preferably before the 2nd week of July. Late sowing causes considerable reduction in yield due to photoperiodicity and excessive soil moisture stress which coincides with the reproductive growth (Chandra et a l . 1 9 8 3 ) . The postrainy season sowing in India should be done in September. In sowings later than 15 October, yields drastically decline (Narayanan and Sheldrake 1 9 7 9 ) . K a u l a n d S e k h o n (1975) c o n c l u d e d t h a t g r a i n y i e l d a n d y i e l d attributing characters were significantly influenced by date of sowing. Sowings on June 1 and June 15 were significantly superior over June 30 and July 15. Significant reduction in grain yield with delay in sowing beyond June 15 was due to reduction in plant height, pods plant-1 and total dry matter production. However delayed sowing gave a high harvest index due to a sharp decline in dry matter production. 20 HRDP SDS no. 9 the In eastern Africa, pigeonpea are sown onset of short rains (Chauhan 1 9 9 0 ) . An optimal seeding depth Ashley 1975; Tayo 1 9 8 3 ) . for pigeonpea in October/November, was 4-5 cm (Khan at and Methods of Sowing and Spacing Three systems of sowings are practiced for pigeonpea. The common is flat sowing, the other methods are broadbed-and-furrow for extra-early group and ridge-and-furrow for the late maturity group. The latter two methods are useful in fields with poor surface drainage and water logging. The raised beds or ridges also provide better aeration and nodulation in comparison to the flat sown crop. Experiments at the Indian Agricultural Research Institute, New Delhi, have shown that pigeonpea sown on a ridge-and-furrow system in fields prone to water logging gave 3 0 % m o r e y i e l d c o m p a r e d t o f l a t s o w i n g (IARI 1 9 7 1 ) . At ICRISAT a broadbed-and-furrow system is used for sowing extra-early g e n o t y p e s , and ridges-and-furrows are used for m e d i u m - and l a t e duration genotypes. Plant spacing depends on maturity duration and cropping system. Extra-early pigeonpea are grown 30 cm between rows with about 10 cm space between plants. The intercropped spacings (Table 4) d e p e n d s on t h e t y p e of intercrop and time to m a t u r i t y (days) o f t h e g e n o t y p e . In traditional production systems, the plant density could be less than 5 p l a n t s m-2 (Pathak 1 9 7 0 ) . These crops mature on residual moisture stored in the soil profile. Natarajan and W i l l e y (1980) f o u n d t h a t 10 p l a n t s m-2 w a s h i g h e r y i e l d i n g t h a n 5 plants m-2. Seed Rate and Seed Treatment The seeding rate of pigeonpea depends on the desired plant d e n s i t y for a g e n o t y p e (early, m e d i u m or l a t e ) , c r o p p i n g system (pure c r o p , m i x e d c r o p , o r i n t e r c r o p ) , g e r m i n a t i o n r a t e o f s e e d , and mass of seed. The plant density recommended for extra-early solecrop pigeonpea is 330 000 plants ha-1, whereas, for early- and mediumm a t u r i t y v a r i e t i e s it ranges from 80 000 to 100 000 p l a n t s ha-1. The plant density for tall and highly branched late varieties could be 50 000-60 000 ha-1. The calculation for seed requirement is given in MP 1. Good quality and pure seed (registered or certified) of the selected variety should be used from a reliable source. Before s o w i n g , t h e s e e d s h o u l d b e t r e a t e d w i t h t h i r a m @ 1.5 g k g - 1 o f s e e d p l u s p e n t a - c h l o r o - n i t r o b e n z e n e (PCNB) [ B r a s s i c o l ® a . i . 1.5 g kg-1] s e e d . This will avoid seed rot and seedling blight diseases (Chandra e t . a l . 1 9 8 3 ) . At ICRISAT we recommend seed HRDP SDS no. 9 21 treatment with 3 g thiram kg - 1 + 3 g carbendazim kg - 1 seed. Weed Management Pigeonpea is a slow-growing crop and mostly cultivated during the rainy season. The crop suffers from early weed infestation. Therefore, it is necessary to keep the crop weed-free during the e a r l y g r o w t h p e r i o d (4-6 w e e k s ) . The weeds can be controlled mechanically or with chemicals. A combination of chemical and mechanical control is more economical. At ICRISAT Center, it was observed that a preemergence application of prometryn (Gesgard® o r C a p a r o l ® a . i . 1.25 k g ha-1) e f f e c t i v e l y c o n t r o l l e d t h e i n i t i a l weeds. A h a n d w e e d i n g 3-4 w e e k a f t e r s o w i n g i s r e q u i r e d t o remove the late emerging weeds. When herbicide is not applied, t w o or t h r e e h a n d w e e d i n g s are r e q u i r e d from the 1st to 6th w e e k of crop growth. Later the crop will be able to smother the weeds (Chauhan et a l . 1 9 8 8 ) . At ICRISAT, hand weedings were always found superior to herbicides, but the herbicides were more economical, hence p r e f e r r e d (Chauhan 1 9 9 0 ) . The other p r e e m e r g e n c e herbicides effective for pigeonpea are p e n d i m e t h a l i n (Stomp® a . i . 1.0-1.5 kg ha-1) or m e t a c h l o r e (Dual® a . i . 1 kg ha-1) . The post-emergence herbicides recommended are flausifop-P b u t y l ( F u s i l a t e ® a . i . 0.2-0.4 k g ha-1) o r b e n t a z o n ( B a s a g r a n ® a . i . 1.0 k g h a - 1 ) a t 2 - 4 l e a f s t a g e (A. R a m a k r i s h n a , 1 9 9 2 , ICRISAT, personal communication). Irrigation Pigeonpea is largely grown as a rainfed crop, however, it is well established that flower initiation and pod setting stages are the most crucial to drought stress. Therefore, irrigation at these stages usually helps the crop (Chandra et a l . 1 9 8 3 ) . The drought-stress symptoms on pigeonpea are indicated by the leaves pointing towards the sun at noon (Chauhan et a l . 1 9 8 8 ) . S i n h a (1981) r e p o r t e d t h e e f f e c t s o f i r r i g a t i o n o n e a r l y pigeonpea and found it difficult to define the adequate water requirement in relation to yield. When irrigation was applied after flowering and at the pod filling there was an increase in biomass in short-, medium-, and long-duration varieties. However, increase in grain yield and biomass did not follow the same pattern. Thus only vegetative growth was increased when water was available, but beyond a certain limit, dry matter was not increased with an increasing water supply. Excessive moisture is detrimental to pigeonpea. It promotes vegetative growth and enhances the i n c i d e n c e o f Phytophthora a n d Alternaria b l i g h t . Therefore, irrigation should be given only when the crop experiences drought stress after flowering and at the pod filling stage (Chandra et a l . 1 9 8 3 ) . Responses to irrigation are more consistent in postrainy season sown pigeonpea. This crop relies on moisture stored in the soil profile. Two or three irrigations, 1 month after 22 HRDP SDS no. 9 sowing, increased seed yield by about 150-160% over the n o n i r r i g a t e d c o n t r o l a t I C R I S A T C e n t e r (Rao e t a l . 1 9 8 3 ) . Harvesting and Threshing Pigeonpea should be harvested when 75-80% of the pods are at physiological maturity. Vanangamudi et al. (1986) s u g g e s t e d harvesting of pigeonpea, about 25 days after anthesis when pods turn brown and are dry. Delayed harvesting, during bad weather, may increase the risk of damage to mature seed. However, B a l a k r i s h n a n a n d N a t a r a j a r a t n a m (1988) suggested harvesting after the 42nd day of anthesis when the dry mass of the seed is low and moisture content is between 20 and 2 4 % . The mature pods could be identified as they are brown and have a dry testa. Traditionally pigeonpea plants are harvested by cutting the stem at the base, with an axe or sickle. The harvested plants are tied in bundles and transported to a threshing floor. These are stacked in upright bundles to dry. The pods and grain are separated by beating the dry plants with sticks or by using a thresher. Another way to harvest pigeonpea is by hand picking the mature pods. This allows the crop to flower and p o d for a second or sometimes a third harvest. Hand picking may not be economical beyond a second flush. When hand picking of pods is not f e a s i b l e , t h e u p p e r b r a n c h e s w i t h m a t u r e p o d s are cut (good for determinate types). Care should be taken in leaving as much of the foliage as possible while removing the pods and allowing the plant to regrow and promote another flush of p o d s . However, this method delays the second harvest and usually results in a lower yield than hand picking (Chauhan et a l . 1 9 8 8 ) . At ICRISAT, pigeonpea is harvested when over 80% of the pods become brown. The harvested plants are left in the field a week for sun drying and threshed in a small plot thresher. The seeds a r e c l e a n e d a n d d r i e d e i t h e r i n a d r i e r a t 35°-43°C o r i n t h e s u n to 6-8% m o i s t u r e c o n t e n t b e f o r e final r e c o r d i n g of y i e l d . Recording Observations Recording of observations depends on the objectives of the experiment. Some important observations for recording in p i g e o n p e a a r e d i s c u s s e d i n M a n a g e m e n t P r o c e d u r e s (MP 2 ) . HRDP SDS no. 9 23 MP 1. Determination of Seed Rate for a Given Plant Density Seed rate ha-1 depends on plant spacing, seed m a s s , and germination percentage of the seed sample. Further, it may be desirable to sow 15-20% more seed than the required to ensure an adequate plant population. Example. Suppose the required plant density is 333 000 plants ha-1, 100-seed m a s s of the variety is 10 g and germination is 90%. T h e s e e d r e q u i r e m e n t ha-1 a n d a p l o t of 15 m2 (3 m x 5 m) w i t h e i g h t rows of 5 m l e n g t h a n d 0.375 m b e t w e e n rows can be calculated. Calculations: a. The seed required 100 seeds 0.90 b. 100 111 = germinable at least 111 seed seed lot was 9 0 % . The seeds 333 Mass required for 333 000 0 0 0 s e e d s ha-1 x 100 seeds 111 seeds (kg) To sow of 20% 37 e. The = f. The h. kg seed 44.4 369 630 seeds g additional ha-1 x 1.20 required m = - 2 requirement m-2 x 15 m2 @ a = HRDP should be for kg 630 100 or seed seeds ha-1 will kg ha-1 will be 37 requirement k g ha-1 = for plot row-1 lot be kg-1 plot-1 g = 36.9 = 4.44 (15 66.6 4.44 = 8.32 24 10 44.4 w 369 g g g m2) g m-2 m-2 will or The seed requirement row-1 will be (The a r e a of r o w is 5 m x 0 . 3 7 5 m = Seed requirement this ha-1 plants seed the ka h a - 1 x 1000 1 0 , 0 0 0 m2 ha-1 seed = 4.44 g. for seeds 3 6 9 6 3 0 s e e d s x 10 a 1 0 0 0 g kg - 1 x 1 0 0 s e e d s d. seed So to obtain 100 seedlings sown as germination of the = c. for 67 1.875 x row-1. SDS no. 9 1.875 be plot-1 g m2 row-1) m-2 row-1 be is Table 6. Pigeonpea seed requirement (kg ha-1) at different densities, germination percentages, and 100-seed mass. Plant 333 100-seed mass Germination 80 (g) 7.0 8.0 29.1 33.3 37.5 9.0 10.0 41.6 11.0 45.8 12.0 13.0 49.9 54.1 14.0 58.3 (Note: for 20% Extra extra 33.3 36.9 40.7 44.4 48.1 51.7 seed seed 80 90 8.8 10.0 7.8 8.9 10.0 11.1 100 25.9 29.7 23.3 26.6 29.9 000 60 Germination (%) 90 ha-1 density 100 000 11.3 12.5 plant (%) 100 7.0 8.0 9.0 000 Germination 80 90 100 5.3 6.0 6.8 7.5 4.7 5.3 6.0 6.7 4.2 4.8 5.4 6.0 36.6 13.8 12.2 10.0 11.0 39.9 43.3 15.0 16.3 13.3 14.4 12.0 13.0 8.3 9.0 9.8 7.3 8.0 8.7 6.6 7.2 7.8 46.6 17.4 15.5 14.0 10.5 9.3 8.4 33.3 (20%) not multiply HRDP included by in the above (%) calculation, 1.2). SDS no. 9 25 MP 2. Recording Morphological Characters Days to emergence. Days are counted from date (irrigation, rain, or adequate moisture) until seedlings have emerged. of sowing 80% of the Plant type. It depends upon the number of primary and secondary branches and angle of branches on the stem. It ranges from upright-compact to spreading type. a. Compact types: the stem. Few branches b. Spreading: c. Semispreading: This and spreading types. Many Days to 50% flowering. of the plants in a plot branches is Days have borne (Fig. narrow angles and broad plant from date of at least one to canopy. intermediate between Days to 75% maturity. This is days after of the pods are physiologically mature. Flowaring pattarn at compact emergence until open flower. 50% emergence 75% until 3). a. Nondatarminata (NDT): The inflorescence develop as axillary racemes from all over the branches, and f l o w e r i n g p r o c e e d s a c r o p e t a l l y (from b a s e t o a p e x ) b o t h within the racemes and on the branches. b. Datarminata (DT): The apical buds of the main shoot and branches develop into inflorescence. Inflorescence a r e p r o d u c e d b a s i p e t a l l y (apex t o b a s e ) , t h e y a r e s h o r t statured and bear clusters of pods at the top. c. Samidatarminata behind the apex basipetally. (SDT): Flowering starts at nodes and proceeds both acropetally and Plant height (cm). Height is recorded at maturity from the b a s e o f t h e p l a n t t o t h e t o p (five p l a n t s are a v e r a g e d ) . Another way to take height across the plot is to put a marked p o l e (cm) a n d n o t e t h e h e i g h t a t t h e c e n t e r o f p l o t . Primary branchas shoots. (number). Secondary branchas branches. (number). Branches emerging Branches borne Number of pods plant-1. Pods are counted at randomly selected plants and the average is 26 HRDP SDS no. 9 from main on primary harvest on recorded. five Number of seeds are counted and 100-seed mass. each treatment pod-1. Seeds in 10 randomly the average is recorded. selected pods One-hundred seeds from the bulk harvest are weighed on an electronic balance. of For genetic studies of crosses and S e e d y i e l d p l a n t - 1 (g) . segregating populations the individual plant yield of seed is recorded on each treatment. Net-plot yield (kg). It pods harvested, threshed, moisture content. is recorded from the net plot for cleaned, dried, and weighed at 7-8% Yield (kg h a - 1 ) . T h e n e t - p l o t y i e l d s a r e c o n v e r t e d t o k g h a - 1 ignoring decimals and rounding to the nearest 10 kg. For e x a m p l e , a y i e l d of 1986 kg ha-1 w i l l be taken as 1990 kg ha-1 and a y i e l d 1984 kg ha-1 will be considered as 1980 kg ha-1. Biological yield. The biological stem, p o d s , seed, and leaves. Shelling as: ratio or seed to husk Seed Harvest index (%) = yield ratio includes (%). This mass is of expressed v i e l d (g) x 1 0 0 P o d y i e l d (g) S e e d y i e l d (g) x 1 0 0 B i o l o g i c a l y i e l d (g) Seed color. P i g e o n p e a m a i n l y h a s o r a n g e s e e d (over 5 0 % ) . I t varies from red, brown, light-brown to w h i t e . The lightcolored seeds are generally bright-green at the filling stage, hence are preferred for vegetable purposes. Seed shape. The most common seed shape is oval. It ranges from elongate, globular to more square. The globular (pea-shaped) trait is preferred for vegetable purposes. D r y s t a l k w e i g h t (kg h a - 1 ) . of seed, pods, and leaves, their economic value. HRDP After threshing and separation the stalks are weighed to estimate SDS no. 9 27 References Aiyer, A.K.Y.N. of Agricultural 1949. Mixed cropping Sciences 19:439-543. in India. Indian Journal Bahadur, B., Rao, M.M., and Rao, K.L. 1981. Studies on dimorphic stamens and pollen (SEM) and its possible role in pollination biology of Cajanus cajan M i l l s p . Indian Journal Botany 4:122-129. of B a l a k r i s h n a n , K., and Natrajaratnam, M. 1988. Identification stable physiological traits for certain cultivars of pigeonpea dry matter accumulation (DMA). Madras Agricultural Journal 73(1):36-41. of in Baldev, B. 1988. Origin, distribution, taxonomy, and morphology. Pages 3-51 in P u l s e crops (Baldev, B., Ramanujam, S., and Jain, H.K., e d s . ) . New Delhi, India: Oxford and IBH Publishing Co. Bisen, S.S., and Sheldrake, A.R. 1981. The anatomy of the pigeonpea. Research Bulletin n o . 5. Patancheru, A . P . 502 324, India: International Crops Research Institute for the Semi-Arid Tropics. 24 pp. Chandra, S., A s t h a n a , A . N . , A l l , M., Sachan, J.N., Lal, S.S., Singh, R.A. and Gangal, L.K. 1983. Pigeonpea cultivation in India - a p a c k a g e of p r a c t i c e s . Kanpur, Uttar Pradesh, India: Directorate of Pulses. 44 pp. Chaudhary, arhar is Chauhan, Y.S. and (In pattern rooting P.A., S.L., adequately eds.). Bhatia, P.C. fertilized. press). of New The tropical Delhi, 1971. Indian root crops India: system (Abdul Tata Profits Farming of when pigeonpea. Salam, McGraw triple 20:27-30. M. Hill and In Wahid, Publishing Company. Chauhan, Y.S. Pages 257-278 Sheila, V.K., 1990. Pigeonpea: optimum agronomic management. in The pigeonpea (Nene, Y.L., Hall, S.D., and eds.) Wallingford, UK: CAB International. Chauhan, Y.S., Johansen, C., and Lateef, S.S. 1988. Package of practices for multiple-harvest system using the short duration pigeonpea cultivar ICPL 87 (Pragati) in Peninsular India. Patancheru, A.P. 502 324, India: International Crops Research Institute for the Semi-Arid Tropics. (Limited distribution.) Dalai, R.C. production Tropical 1980. by Agriculture Dalal, R.C., on nutrition 69:854-857. 28 field and and Nutrient grown element pigeonpea (Trinidad) concentration [Cajanus and (L.) dry matter Millsp.], 57 (2):107-112; Quilt, P. 1977. Effects grain yield of pigeonpea. HRDP cajan of N, P, liming and Mo Agronomy Journal SDS no. 9 De, D.N. 1974. Pigeonpea. Pages 79-87 in Evolutionary studies in world crops: diversity and change in the Indian subcontinent ( H u t c h i n s o n , J., e d . ) . London, UK: Cambridge University Press. Edwards, D.G. 1981. Development of research on pigeonpea nutrition. Pages 205-211 in Proceedings of the International Workshop on Pigeonpeas, 15-19 Dec 1980, ICRISAT Center, India. Vol. 1. Patancheru, A.P. 502 324, India: International Crops Research Institute for the Semi-Arid Tropics. FAO 1991. Pigeonpeas. Nations. Topics on Food and Agricultural Agricultural Statistics (Agro Stat) 0197 Organization of the United Green, J.M., Sharma, D., Saxena, K.B., Reddy, L.J., and Gupta, S.C. 1979. Pigeonpea breeding at ICRISAT. Presented at the Regional Workshop on Tropical Grain Legumes, 18-22 Jun 1979, University of West Indies, St. Augustine, Trinidad. (Limited distribution.) IARI (Indian Agricultural Research Institute). in pulse production. New Delhi, India: IARI. 1971. New ICAR (Indian Council of Agricultural R e s e a r c h ) . 1979. progress reports of the All India Coordinated Agronomic Project for 1972-73 to 1978. New Delhi, India: ICAR. Johansen, C. Katyal, J.C. 221-226 in Pigeonpea: mineral nutrition. Pages (Nene, Y.L., Hall, S.D., and Sheila,V.K., CAB International. 1981. Pigeonpeas, 15-19 A.P. for Kaul, J.H., dates of Agronomy the and Dec 502 in T.N., and ICRISAT India: and Center, in pigeonpea. Workshop India. International Pages on Vol.2. Crops Research Tropics. H.S. 1975. phosphorus Response levels. of pigeonpea Indian Journal to of Proceedings of ICRISAT An approach to the breeding of pigeonpea Millsp.]; formation of composites. Euphytica Ashley, J.M. Experimental K.R., in India: 1980, research International 209-231 eds.). 20 (4):376-377. fertilizers 1980, the 324, Sekhon, planting pigeonpea. Kulkarni, of Semi-Arid Khan, T.N. 1973. [ C a j a n u s cajan (L.) 22: 373-377. Khan, Micronutrient Proceedings Patancheru, Institute Annual Research 1990. in The pigeonpea Wallingford, UK: vistas and Panwar, India: a K.S. critical International Center, International 1975. India. Crops Factors Agriculture 11: 1981. Response review. Workshop Vol. Research 1. affecting on plant stand 315-322. Pages of Pigeonpeas, Patancheru, Institute pigeonpea 212-220 for 15-19 A.P. the to in 502 Dec 324, Semi-Arid Tropics. HRDP SDS no. 9 29 Kumar Rao, J.V.D.K. 1990. Pigeonpea: nitrogen fixation. Pages 233-256 in The pigeonpea (Nene, Y.L., Hall, S.D., and Sheila, V.K., eds.). Wallingford, UK: CAB International. Mahta, D.N., and Dave, B.B. 1931. Studies in Memoirs of the Department of Agriculture in India Series) 19:1-25. Cajanus indicus. (Botanical Narayanan, A., and Sheldrake, A.R. 1979. Pigeonpea (Cajanus cajan) as a w i n t e r c r o p in P e n i n s u l a r India. Experimental Agriculture 15:91-95. Natarajan, M., and Willey, R.W. 1980. Sorghum-pigeonpea inter cropping and the effect of plant population density. 1. Growth and yield. Journal of Agricultural Science (UK) 95:51-58. Nichols, R. 1964. Studies on the major element deficiencies the pigeonpea ( C a j a n u s cajan) in sand culture. 1. Foliar symptoms of the major-element deficiencies. Plant and Soil 21(3): 377-387. of Nichols, R. 1965. Studies on the major element deficiencies of the pigeonpea ( C a j a n u s cajan) in sand culture. 2. The effects of major element deficiencies on nodulation, growth, and mineral composition. Plant and Soil 22(1):112-126. Panwar, K.S. agronomy Annual on All Kharif Jun 1975, 1975. Report (1974-1975) India on CSAUT, Coordinated Pulses, New at Indian Delhi, the work Kanpur. Pulses on kharif Improvement Agricultural India. done Presented (Limited Research at pulses the Project 11th Workshop Institute, 9-12 distribution) Panwar, K.S., and Misra, A.S. 1973. Recommendations for the agronomy of kharif pulses. Presented at the Ninth Annual All India Coordinated Pulses Improvement Project Workshop on Kharif Pulses, Indian Agricultural Research Institute, 7-9 J u n 1 9 7 3 , New Delhi, India. (Limited distribution) Pathak, G.N. 1970. Redgram. Pages 14-53 in Pulse crops India. New Delhi, India: Indian Council of Agricultural Research. Plukenet, L. 1692. Phytographia 3, Rao, M.K.V., and Rao, R.G. 1974. developing and germinating seeds of Plant Physiology 17 (1/2):65-72. Rao, 1983. I.M., Vankataratnam, Response International Raddy, L.J. pigeonpea 30 1990. UK: N., irrigation Pigeonpea (Nene, Wallingford, to Pigeonpea: Y.L., CAB Hall, D.G., postrainy Newsletter and 3. like substance in Indian Journal of Shaldraka, season and Pages Sheila, International. HRDP Figure A.R. pigeonpea. 2:35-36. morphology. S.D., 213, Gibberellin pigeonpea. Faris, in Table of SDS no. 9 47-88 V.K., in The eds.). Reddy, S.J., and Virmani, S.M. 1981. Pigeonpea and its climatic environment. Pages 259-270 in Proceedings of the International Workshop on Pigeonpeas, 15-19 Dec 1980, ICRISAT Center, India. Vol. 1. Patancheru, A.P. 502 324, India: International Crops Research Institute for the Semi- Arid Tropics. R e m a n a n d a n , P., Sastry, D.V.S.S.R., and Mengesha, M . H . 1988. ICRISAT pigeonpea germplasm catalogue: evaluation and analysis. Patancheru, A.P. 502 324, India: International Crops Research Institute for the Semi-Arid Tropics. 95 pp. Saxena, M.C., and Singh, Y. 1970. Relative susceptibility of important varieties of some pulses and soybean to zinc deficiency. Presented at the Third Annual Workshop of Coordinated Scheme on Micro-nutrients of soils, 5-7 O c t 1 9 7 0 , Ludhiana, Punjab, India. Sehgal, C.B., Gandhi, V., and Varma, B. 1987. Histological and histochemical studies on the cotyledons of some legumes. II. Reserve metabolites during seed development. Cytologia 52: 847-858. Sharma, D., and Hybridization H., Crop eds.). Green, J.M. crop plants of Madison, Science WI, Society 1980. Pigeonpea. (Fehr, Walter USA: American R. Pages and Society 471-481 Hadley, of in Henry Agronomy and America. of S h a r m a , D . , R e d d y , L.J., G r e e n , J.M., and Jain, K.C. 1981. International adaptation of pigeonpeas. Pages 71-80 in Proceedings of the International Workshop on Pigeonpeas, 15-19 Dec 1980, ICRISAT Center, India. Vol. 1. Patancheru, A.P. 502 324, India: International Semi-Arid Tropics. Crops Research Institute for the Sheldrake, A.R. 1984. Pigeonpea. Pages 385-417 in The physiology of tropical field crops (Goldsworth, P.R., and N.M., e d s . ) . Chichester, USA: John Wiley & Co. Sheldrake, nutrient A.R., Agricultural Shetty, and uptake in Sciences S.V.R., and sorghum/pigeonpea some International Sinha, S.K. pigeonpea. Workshop Vol. Rao, 1. Research on Workshop Water 283-288 Pigeonpeas, Patancheru, Institute 1979. 238-248 the HRDP Weed in development Journal management and of A.P. 502 Institute 502 1980, 324, the ICRISAT India: Semi-Arid Jan in - the 1979, Semi-Arid grain of of systems India: the and Proceedings Dec 10-13 324, for studies intercrop Proceedings availability in 15-19 A.P. for Growth cajan). Inter-cropping, Research 1981. 1979. millet/groundnut Patancheru, Crops Pages A.N. on A. (Cajanus 92:513-526. pearl Pages India. International (UK) and observations. Hyderabad, Narayanan, pigeonpea Fisher, yield Tropics. in International Center, International India. Crops Tropics. SDS no. 9 31 Sundaraj, D.D., New Delhi. The and Thulasidas, G. 1980. Macmillan Company of India Botany of field Ltd. 4 96 pp. crops. Tayo, T.O. 1983. Growth, development, and yield of pigeonpea. ( C a j a n u s cajan (L.) Millsp.) in the low land tropics 4. Effect of sowing depth. Journal of Agricultural Science, Cambridge. 101:435-440. Tandon, H.L.S. 1987. Phosphorus research and agricultural production in India. Fertilizer Development and Consultation Organization, New Delhi, India. 104 pp. van der Maesen, L.J.G. 1980. India is the native home of pigeonpea. Pages 257-262 in Libergratulatorius in honorem de Wit. Landbouwhogeschool (Arends, J.C., B o e l e m a , G., d e C.T., and Leeuwenberg, A.J.M., e d s . ) . Miscellaneous Paper 19. Wageningen, N e t h e r l a n d s : H. Veenman and B.V. Zonen. the H.C.D. Groot, no. Vanangamudi, K., Karivaratharaju, T.V., Balakrishna, K., M a l l i k a and Natarajaratnam, N. 1986. Effect of stage of harvest on seed quality in pigeonpea cultivars. International Pigeonpea Newsletter 5:31-32. Vavilov, N.I. breeding of Vernon Royes, crop plants 1951. cultivated W. 1976. (Simmonds, The origin, plants. variation, Chronica Pigeonpea. N.W., e d . ) . immunity Botanica Pages 154-156 London, UK: and 13(1-6):1-366. in Evolution of Longmans. Willay, R.W., Rao, M.R., and Natarajan, M. 1981. Traditional cropping systems with pigeonpea and their improvement. Pages 11-25 in Proceedings of the International Workshop on Pigeonpeas, 15-19 Dec 1980, ICRISAT Center, India. Vol. 1. Patancheru, A.P. 502 324, India: International Crops Research Institute for the Semi-Arid Tropics. Zeven, A.C., and Zhukovsky, P.M. plants. Pages 549-566 in The new London, UK: Oxford University. 32 HRDP 1975. Dictionary of cultivated systematics (Huxby, J., e d . ) . SDS no. 9 Evaluation Select at the 1. The the most appropriate end of the booklet. pigeonpea belongs a) Atylosia. c) Arachis. answer to 2. The word pigeonpea a) USA. c) Zambia. first 3. The primary center a) Africa. c) Asia. of 4. The average productivity a ) 2 0 0 kg h a - 1 . c) 7 6 0 k g h a - 1 . 5. Pigeonpea have a) adventitious roots. c) seminal roots. the genus b) d) check the correct answer Cajanus. Cicer. originated from b) Barbados, d) India. origin pigeonpea and of of pigeonpea is b) Europe. d) Australia. pigeonpea is b) 500 kg ha-1. d) 900 kg ha-1. b) d) staminal roots. tap root. 6. The extensive a r e a is to a a) 2 0 cm c) 60 c m 7. Nodulations in pigeonpea a) meliloti group. c) japonicum group. 8. The greatest number of nodules in during the a) seedling stage. b) vegetative phase. c) early reproductive phase. d) maturity stage. 9. The branching pattern that first and then enters into a) nondeterminate. c) semideterminate. 10. When flowering starts at nodes behind the apex and proceeds both acropetally and basipetally it is a) nondeterminate. b) determinate. c) semideterminate. d) none of the above. HRDP root development depth. b) 40 cm d) 80 cm are by in the upper rhizobia belonging b) trifolii group. d) cowpea group. pigeonpea are to soil the observed completes the vegetative phase the reproductive phase is the b) determinate. d) none of the above. SDS no. 9 33 11. Late-maturing pigeonpea genotypes start branching from different angles, and when most of the pods are at the top, the plants are a) nondeterminate. b) determinate. c) semideterminate. d) none of the above. 12. The The flowering on pigeonpea, branches proceeds a) basipetally. c) determinate. 13. 14. 15. 16. first two leaves in pigeonpea seedlings are a) pinnately trifoliate. b) simple and opposite. c) c o m p o u n d . d) b o t h a a n d c. The The pigeonpea flowers are a) incomplete. b) complete, bisexual, c) monoecious. d) dioecious. calyx is a) polysepalous. b) within b) d) raceme and on zygomorphic. gamosepalous. petal with b) keel. the c) standard. obliquely a) wing. 18. An obtuse (round), a) wing. 19. The aestivation of the pigeonpea inflorescence is a) ascending imbricate. b) descending imbricate. c) free. Pigeonpea anthers are a) monoadelphous. Anthesis in pigeonpea a) 0800-1200. c) 0900-1300. 22. The 24. 34 In pigeonpea a) 1 w e e k . c) 3 w e e k s . Pigeonpea seed flowering. a) 20 days c) 40 days claw is the standard. incurved, and boat-shaped petal is b) keel. c) standard. 21. 23. incurved c) free. An peak anthesis a) 0800-0900. c) 0900-1000. an is c) 17. 20. after 10 b) diadelphous. takes place from b) 0600-1600. d) 1000-1800. recorded b) d) in pigeonpea 0700-0800. 1100-1200. pollination, the pod is b) 2 w e e k s . d) 4 w e e k s . attains physiological b) d) HRDP the and usually period the acropetally. nondeterminate. The broad, large, and auricled petal a) wing. b) keel. obovate the maturity 30 days 50 days SDS no. 9 is from visible in in after 25. The deep constricted a) beaded. c) shrunken. 26. The germination a) epigeal. 27. 28. The in application of weeks before a ) 1-2 b ) 2-4 pod of pigeonpea pigeonpea is b) flattish. d) shrivelled. is b) organic manure sowing. c ) 5-6 d) To increase the pigeonpea necessary to apply a) organic manure. c) l i m e . hypogeal. in should be 8-10 production b) d) pigeonpea in acidic soil it is single superphosphate. gypsum. 29. The recommended rate production is a) 5-10 b) 1 2 - 1 6 30. Uniform pale-green young leaves and interveinal older leaves are deficiency symptoms of a) phosphorus. b) potassium. c) n i t r o g e n . d) zinc. chlorosis 31. Small stunted plants deficiency of a) potassium. c) n i t r o g e n . due 32. to apply phosphorus kg ha-1. c) 17-26 d) 30-35 with dark-green b) d) Appearance of light-green patches around the leaf margin which turn symptoms of a) potassium. b) c) nitrogen. d) (P) leaves for are pigeonpea to the phosphorus. calcium. in interveinal areas or brown are the deficiency phosphorus. calcium. 33. Yellowing on leaf tips followed by scorching and plants are the main deficiency symptoms of a) potassium. b) phosphorus. c) nitrogen. d) calcium. 34. Stunted growth, narrowing of leaves with pale-green appearance, interveinal chlorosis starting from tip and spreading to other area of leaves except the midrib. The curling and shedding of leaves is the deficiency symptoms a) phosphorus. b) potassium. c) zinc. d) iron. 35. stunting When interveinal areas of young leaves become pale-green veins and adjacent tissues remain dark-green, it is the deficiency symptoms of a) zinc. b) nitrogen. c) iron. d) calcium. HRDP of SDS no. 9 of of but 35 36. Fading of the lamina of the middle leaves without affecting the veins with small white and brown spots, and brown necrotic lesions with a reduction of leaf number, area and size, late flowering and the arrest of the apical shoot growth are deficiency symptoms of a) zinc. b) iron. c) calcium. d) manganese. 37. Formation of dieback, rosetting, multiple branches, of seedlings are deficiency symptoms of a) iron. b) manganese. c) zinc. d) boron. 38. An 39. An economic dose of fertilizer are kg ha-1. a) 10-20-30 c) 20-20-50 important a) iron. c) zinc. element in and death nitrogen fixation is b) boron. d) molybdenum. for p i g e o n p e a b) d) with N, P, A c o m b i n a t i o n of crops in space and time to p r o v i d e a and stable level of returns is a) a crop r o t a t i o n . b) crop m a n a g e m e n t . c) a c r o p s e q u e n c e . d) a c r o p p i n g s y s t e m . 41. The a) c) suitable genotypes late-maturing types. early-maturity types, of b) d) Postrainy season pigeonpea a) a m i x e d c r o p . c) a c a s h c r o p . 43. The best time is in t h e 1st a) May. b) 44. An 45. Late sowing of pigeonpea a) yield. c) root m a s s . 46. The 47. A b r o a d b e d - a n d - f u r r o w system of for a) early-maturing pigeonpea. c) late-maturing pigeonpea. 36 optimal depth a) 2-3 c m . c ) 7-8 c m . for pigeonpea for high monocropping are medium-maturity types. extra-early types. 42. of sowing fortnight June. c) K 20-17-17 30-40-60 40. most and are cultivated as b) an i n t e r c r o p . d) a m o n o c r o p . of extra early pigeonpea of July. d) August. pigeonpea at ICRISAT sowing is b ) 4-5 c m . d ) 9-10 c m . increases b) biomass. d) harvest index. most common practice of pigeonpea cultivation is on a) broadbed-and-furrows. b) ridges-and-furrows. c) bunds. d) flat b e d s . HRDP cultivation b) d) is most suitable medium-maturing pigeonpea. extra-early-maturing types SDS no. 9 48. The 49. The plant population varieties is a) 80 000 h a - 1 . c) 600 000 h a - 1 . row and plant spacing a) 50 a n d 40 cm. c) 75 a n d 20 cm. in extra-early pigeonpea b) 40 a n d 60 cm. d) 30 a n d 10 cm. recommended for late-maturing To p r e v e n t seed rot and seedling blight diseases, recommended to treat the seed with a) thiram. b) carbendazim. c) thiram+carbendazin. d) penta-chloro-nitro-benzene (PCNB). 51. For an effective weed control it herbicides. a) presowing b) preemergence for weed is necessary c) to control in pigeonpea b) alachlor. d) glyphosate. 53. When preemergence herbicide is applied, the first weeding is required weeks after sowing, a ) 1-2 b ) 3-4 c ) 6-7 d ) 8-9 54. At noon when leaves are pointing towards the symptoms of a) nitrogen, b) phosphorus, c) drought. 55. The 56. Pigeonpea should be harvested when mature. a) 40-50 b) 50-60 c) 70-80 critical periods for drought stress a) seedling and vegetative stages. b) flowering and seedling stages. c) flowering and pod-filling stages. d) maturity and flowering stages. Before storage it is necessary to moisture content of a) 10-15%. b) 10-16%. c) dry it d) of is stress wilt. the pods are the are 80-90 pigeonpea 8-10%. SDS sun is hand in pigeonpea % d) is postemergence An HRDP it apply 52. 57. pigeonpea 100 000 ha-1. 200 000 ha-1. b) d) 50. important herbicide a) fluchloralin. c) 2-4D. are no. 9 d) seed to a 6-8%. 37 Correct responses to the questions. 1. b ) ; 9. b ) ; 16. c ) ; 23. a ) ; 30. 37. c) ; d); 44. b ) ; 51. b ) ; 38 2. c ) ; 3. c ) ; 10. a ) ; 11. c ) 17 . a ) ; 18. b ) 2 4 . b) ; 25. a ) 4. ; ; ; 31. b ) ; 3 8 . d) ; 45. d ) ; 32. a ) ; 39. b); 46. d ) ; 52. a ) ; 53. b ) ; HRDP c) ; 5. 12. b ) ; 19. b ) ; 26. b ) ; 33. a ) ; 40. d ) ; 47. d ) ; 54. c ) ; d); 13. 20. 27. 34. 41. 6. b); b); c) ; 7. d); 8. c ) ; 15. b ) ; 14. b ) ; 22. c ) ; 21. b ) ; b); 48. c); d); d); 28. c ) ; 35. c ) ; 42. d); 49. a ) ; 55. c); 56. SDS no. 9 c ) ; 29. c ) ; 36. d ) ; 43. b ) ; 50. c ) ; 57. d ) .