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FUNCTIONS OF LEAVES What are leaves Leaves are an essential plant organ because they play a vital part in the nutrition of plants. Leaves vary considerably in shape and form, but they are all have the same function. A leaf is a lateral outgrowth from a stem, arises at a node and possesses a bud in its axil. The leaf has two main parts; the blade (lamina) and the stalk (petiole). When there is no petiole the leaf is said to be sessile. The lamina is supported by a system of veins, and when a vein or midrib with numerous branches forms a network, the venation is said to be reticulate. This pattern is typical of dicotyledons. When several veins of approximately equal size run parallel to each other the venation is said to be parallel. This arrangement is typical of monocotyledons. Leaves may be simple or compound (the latter has more than one blade and the component leaflets are called pinnae). Pinnae may be further divided into pinnules. Leaf arrangement around the stem, the leaf shape, leaf base, leaf apex and leaf margin are all morphological characters used in identification. Functions of leaves Photosynthesis The production of food (sugars) using sunlight, carbon dioxide and water. Transpiration The process by which plants lose water in the form of water vapour. Leaves have wide variations in morphology. In many instances there are adaptations to the particular climatic conditions of the plant's environment, so that an adequate surface for photosynthesis is combined with features to minimize excessive water storage. The thick fleshy bases of leaves that make up most of the Daffodil bulb accumulate large quantities of food. In sweet peas, tendrils are the equivalent of leaflets. External leaf structure Lamina the blade of the leaf. Margin the edge of the leaf. Veins conducting tissue of the leaf. Contains xylem which conducts water and minerals from the roots and phloem which moves sugars and other products from the leaves to the rest of the plant. Midrib the central conducting vein in dicots. Petiole the stalk which holds the leaf. Leaf base point at which the lamina attaches to petiole. Stipule a small leaf-like appendage at the base of the petiole. Node the slightly enlarge portion of the stem where leaves and buds arise, and where branches originate. Internode the region of the stem between two successive nodes. Axil the upper angle between the petiole of a leaf and the stem from which it grows. Axillary bud. an embryonic shoot which lies at the junction of the stem and petiole Apex. is the tip or end farthest from the base or point of attachment How do we tell if we are looking at a leaf or leaflet? The easiest way to determine this is to look at the position of the buds. Buds occur in the axils of leaves, not in the axils of leaflets. Some wattles have phyllodes rather than leaves. The petiole is enlarged and usually flattened and performs the functions of a leaf. Internal leaf structure In most leaves there are three types of tissue. These are as follows: Epidermis On both the upper and lower surface of the leaf. Usually consist of a single layer of tightly fitting cells. It is covered with a waxy coating, the cuticle, that helps to prevent water loss and hence drying out of the leaf. Stomata (singular stoma or stomate) are pores that occur in the epidermis, usually more abundantly in the lower epidermis. Carbon dioxide and oxygen pass in and out through these pores. Water that is evaporated through transpiration also passes out through the stomata. Two guard cells surround each stomate. The guard cells swell and shrink to regulate the opening and closing or the stomate. Hairs may also grow out from the epidermis. Mesophyll tissue The mesophyll tissue is composed of parenchyma cells and forms the two layers between the upper and lower epidermis. The palisade mesophyll cells are tall and cylindrical and contain many chloroplasts (with the pigment chlorophyll) for photosynthesis. The spongy mesophyll cells are below the palisade mesophyll, are irregular in shape, and contain fewer chloroplasts. They have large air spaces between them. Carbon dioxide and oxygen move through this layer. This means that most of the food-producing cells have free access to these gases. Veins or vascular bundles These are the conducting tissues. Xylem occupies the upper part of the veins, phloem the lower part. The veins transport minerals and water (xylem), and products of photosynthesis (phloem).