Download Concept 41.5: Evolutionary adaptations of vertebrate digestive

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The Small Intestine
• The small intestine is the longest section of the
alimentary canal
• It is the major organ of digestion and absorption
LE 41-19
Liver
Bile
Gallbladder
Stomach
Acid chyme
Intestinal
juice
Pancreas
Duodenum of
small intestine
• The pancreas produces proteases, proteindigesting enzymes that are activated after
entering the duodenum
LE 41-20
Pancreas
Membrane-bound
enteropeptidase
Inactive
trypsinogen
Other inactive
proteases
Lumen of duodenum
Trypsin
Active
proteases
• The liver produces bile, which aids in digestion
and absorption of fats
• The epithelial lining of the duodenum, called the
brush border, produces several digestive enzymes
LE 41-21
Carbohydrate digestion
Protein digestion
Nucleic acid digestion
Fat digestion
Oral cavity, Polysaccharides Disaccharides
pharynx,
Salivary amylase
esophagus
Smaller polysaccharides, maltose
Stomach
Proteins
Pepsin
Small polypeptides
Lumen of
small intestine
Polysaccharides
Polypeptides
Pancreatic amylases
Pancreatic trypsin
and chymotrypsin
Maltose and other
disaccharides
DNA, RNA
Pancreatic
nucleases
Nucleotides
Pancreatic carboxypeptidase
Pancreatic lipase
Amino acids
Disaccharidases
Monosaccharides
Bile salts
Fat droplets
Smaller polypeptides
Epithelium
of small
intestine
(brush
border)
Fat globules
Glycerol, fatty
acids, glycerides
Small peptides
Nucleotidases
Dipeptidases, carboxypeptidase, and
aminopeptidase
Nucleosides
Amino acids
Nucleosidases
and phosphatases
Nitrogenous bases,
sugars, phosphates
LE 41-22
Key
Liver
Stimulation
Enterogastrone
Inhibition
Gallbladder
Gastrin
CCK
Stomach
Pancreas
Secretin
Duodenum
CCK
Absorption of Nutrients
• The small intestine has a huge surface area, due
to villi and microvilli that are exposed to the
intestinal lumen
• The enormous microvillar surface greatly
increases the rate of nutrient absorption
LE 41-23
Key
Vein carrying blood
to hepatic portal
vessel
Nutrient
absorption
Microvilli
(brush border)
Blood
capillaries
Epithelial
cells
Muscle layers
Epithelial cells
Large
circular
folds
Villi
Lacteal
Villi
Intestinal wall
Lymph
vessel
Fat globule
• Amino acids and sugars pass
through the epithelium of
the small intestine and enter
the bloodstream
• After glycerol and fatty acids
are absorbed by epithelial
cells, they are recombined
into fats within these cells
Bile salts
Fat droplets
coated with
bile salts
Micelles made
up of fatty acids,
monoglycerides,
and bile salts
Epithelium
of small
intestine
Epithelium
of lacteal
Lacteal
The Large Intestine
• The large intestine, or colon, is connected to
the small intestine
• Its major function is to recover water that has
entered the alimentary canal
• The colon houses strains of the bacterium
Escherichia coli, some of which produce
vitamins
Concept 41.5: Evolutionary adaptations of vertebrate digestive
systems are often associated with diet
• Digestive systems of vertebrates are variations
on a common plan
• However, there are intriguing adaptations, often
related to diet
Some Dental Adaptations
• Dentition, an animal’s assortment of teeth, is one
example of structural variation reflecting diet
• Mammals have specialized dentition that best
enables them to ingest their usual diet
LE 41-26
Incisors
Molars
Canines
Premolars
Carnivore
Herbivore
Omnivore
Stomach and Intestinal Adaptations
• Herbivores generally have longer alimentary canals
than carnivores, reflecting the longer time needed
to digest vegetation
LE 41-27
Small intestine
Stomach
Small
intestine
Cecum
Colon
(large
intestine)
Carnivore
Herbivore
Symbiotic Adaptations
• Many herbivores have fermentation chambers,
where symbiotic microorganisms digest cellulose
• The most elaborate adaptations for an herbivorous
diet have evolved in the animals called ruminants
LE 41-28
Intestine
Rumen
Reticulum
Esophagus
Abomasum
Omasum